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Paper Citation Record · LEDGER

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images

As of 17 August 2026, this Paper Citation Record lists 43 of 43 outbound references and 0 inbound Pith citation observations for arXiv:2412.02601.

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pith.paper-citation-record.v1
2412.02601 v2

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measured 43 of 43 reference resolution

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Pith citing papers itemized under the disclosed page cap.

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Reference resolution

43 of 43 outbound references displayed

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External citation measurements

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Outbound references

Observation 64fc0e29-d7e9-42e4-84f4-589822b8a29a · outbound

This paper cites Principal component analysis.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Principal component analysis

Reference 1

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Observation c3440960-27da-43cf-8847-5af96dafdbca · outbound

This paper cites Goodwin, Anubhav Sinha, et al.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Goodwin, Anubhav Sinha, et al

Reference 2

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Observation 2bdb0740-e903-4220-8fef-d5ab2e09ccd9 · outbound

This paper cites Spatial deconvolution of her2-positive breast tumors reveals novel intercellular relationships.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Spatial deconvolution of her2-positive breast tumors reveals novel intercellular relationships

Reference 3

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Observation 2198fa25-7791-4926-9529-de88f83b4459 · outbound

This paper cites Layer Normalization.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Layer Normalization

Reference 4

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Observation 7b7d4c1b-c84f-44e3-9acd-2b6a8e1ffb6f · outbound

This paper cites Boettiger, Jeffrey R.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Boettiger, Jeffrey R

Reference 5

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Observation 980050ae-fe9b-4490-b717-593d5ae97c9e · outbound

This paper cites Towards a general-purpose foundation model for computational pathol- ogy.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Towards a general-purpose foundation model for computational pathol- ogy

Reference 6

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Observation fb46d516-a3ad-4984-b9d3-fab7a593caa2 · outbound

This paper cites Accurate spatial gene expression prediction by integrating multi-resolution features.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Accurate spatial gene expression prediction by integrating multi-resolution features

Reference 7

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Observation b3a2e4eb-eaef-4d30-9850-45d8c544de12 · outbound

This paper cites Borm, Amit Zeisel, et al.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Borm, Amit Zeisel, et al

Reference 8

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Observation 911083e1-7158-4c2e-a016-7a42a12476d2 · outbound

This paper cites Sys- tematic comparison of single-cell and single-nucleus rna- sequencing methods.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Sys- tematic comparison of single-cell and single-nucleus rna- sequencing methods

Reference 9

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Observation f2f9981f-bcf2-4e31-929b-2946ffb7fe75 · outbound

This paper cites An Image is Worth 16x16 Words: Transformers for Image Recognition at Scale.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images An Image is Worth 16x16 Words: Transformers for Image Recognition at Scale

Reference 10

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Observation ec0d59d1-5b08-4bc2-9992-31ae49357147 · outbound

This paper cites Transcriptome-scale super-resolved imaging in tissues by rna seqfish+.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Transcriptome-scale super-resolved imaging in tissues by rna seqfish+

Reference 11

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Observation 27c5f249-4091-4810-ac0f-1ddb75b838c2 · outbound

This paper cites Inductive representation learning on large graphs.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Inductive representation learning on large graphs

Reference 12

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This paper cites In- tegrating spatial gene expression and breast tumour morphol- ogy via deep learning.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images In- tegrating spatial gene expression and breast tumour morphol- ogy via deep learning

Reference 13

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Observation 5c4f34f3-08f1-46dc-8e37-4e6f775b5650 · outbound

This paper cites Deep residual learning for image recognition.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Deep residual learning for image recognition

Reference 14

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Observation f2040cca-a0ca-4bb6-b65f-20b3da825677 · outbound

This paper cites A systematic evaluation of single-cell rna-sequencing imputation methods.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images A systematic evaluation of single-cell rna-sequencing imputation methods

Reference 15

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This paper cites Densely connected convolutional net- works.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Densely connected convolutional net- works

Reference 16

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Observation 5272d344-baa9-40fc-870b-92ba6365cad6 · outbound

This paper cites Saver: gene expression recovery for single-cell rna sequencing.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Saver: gene expression recovery for single-cell rna sequencing

Reference 17

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Observation 0390b08d-5cc7-4209-92a9-46bd790cf428 · outbound

This paper cites A visual–language foundation model for pathology image anal- ysis using medical twitter.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images A visual–language foundation model for pathology image anal- ysis using medical twitter

Reference 18

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Observation effe6b75-e281-4e97-ad76-148d2302fe06 · outbound

This paper cites Mrna stability and overexpression of fatty acid synthase in human breast cancer cell lines.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Mrna stability and overexpression of fatty acid synthase in human breast cancer cell lines

Reference 19

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Observation 10b02546-8935-4661-ab81-e1e5d798cee9 · outbound

This paper cites HEST-1k: A Dataset for Spatial Transcriptomics and Histology Image Analysis.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images HEST-1k: A Dataset for Spatial Transcriptomics and Histology Image Analysis

Reference 20

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Observation 8256a0d0-e829-4685-a16e-1dd15c090ba4 · outbound

This paper cites Multimodal analysis of composition and spatial architecture in human squamous cell carcinoma.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Multimodal analysis of composition and spatial architecture in human squamous cell carcinoma

Reference 21

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Observation a3f5152f-526f-47e0-86cc-b4f147b952a8 · outbound

This paper cites Thitogene: a deep learning method for predicting spatial transcriptomics from histological images.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Thitogene: a deep learning method for predicting spatial transcriptomics from histological images

Reference 22

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Observation 9bb98f67-6c44-480b-8df9-da2ab7fb2591 · outbound

This paper cites Elevated expression of gnas pro- motes breast cancer cell proliferation and migration via the pi3k/akt/snail1/e-cadherin axis.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Elevated expression of gnas pro- motes breast cancer cell proliferation and migration via the pi3k/akt/snail1/e-cadherin axis

Reference 23

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Observation a0576a09-ab9e-4d1c-8e6f-5cad46365893 · outbound

This paper cites Bayesian approach to single-cell differential expression anal- ysis.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Bayesian approach to single-cell differential expression anal- ysis

Reference 24

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This paper cites Fast, sen- sitive and accurate integration of single-cell data with har- mony.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Fast, sen- sitive and accurate integration of single-cell data with har- mony

Reference 25

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Observation dbf8b0dc-a0ed-4352-8c4e-4f573bb84d0d · outbound

This paper cites SPCS: a spatial and pattern combined smoothing method for spatial transcriptomic expression.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images SPCS: a spatial and pattern combined smoothing method for spatial transcriptomic expression

Reference 26

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This paper cites Visual language pretrained multiple instance zero-shot transfer for histopathology images.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Visual language pretrained multiple instance zero-shot transfer for histopathology images

Reference 27

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Observation 2564051e-1572-43a7-a950-264d2c99f13b · outbound

This paper cites A visual-language foundation model for computational pathol- ogy.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images A visual-language foundation model for computational pathol- ogy

Reference 28

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Observation 3fbed033-e3c0-4e97-a6de-682e29fcb332 · outbound

This paper cites EAGS: efficient and adaptive Gaussian smoothing applied to high-resolved spatial transcriptomics.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images EAGS: efficient and adaptive Gaussian smoothing applied to high-resolved spatial transcriptomics

Reference 29

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Observation 84fb01c5-b4a7-4f63-b3cd-610a9efd6e5d · outbound

This paper cites Leverag- ing information in spatial transcriptomics to predict super- resolution gene expression from histology images in tumors.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Leverag- ing information in spatial transcriptomics to predict super- resolution gene expression from histology images in tumors

Reference 30

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This paper cites St ˚ahl, Fredrik Salm ´en, Sanja Vickovic, et al.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images St ˚ahl, Fredrik Salm ´en, Sanja Vickovic, et al

Reference 31

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This paper cites Patches Are All You Need?.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Patches Are All You Need?

Reference 32

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This paper cites Re- covering gene interactions from single-cell data using data diffusion.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Re- covering gene interactions from single-cell data using data diffusion

Reference 33

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verified fuzzy
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Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.

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Observation c6310ccb-cc82-4941-a34b-121e8148ba3b · outbound

This paper cites Attention is all you need.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Attention is all you need

Reference 34

Resolution
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no resolver link, observed 2026-08-11T23:24:41.481264Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

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Observation e184a329-1eae-4f0f-8baa-666d2b672a9b · outbound

This paper cites Graph Attention Networks.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Graph Attention Networks

Reference 35

Resolution
verified fuzzy
raw_fallback, observed 2026-08-11T23:24:41.780923Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.

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Observation 928f65db-29b0-49d9-87bd-71997bf91568 · outbound

This paper cites High-definition spatial transcriptomics for in situ tissue pro- filing.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images High-definition spatial transcriptomics for in situ tissue pro- filing

Reference 36

Resolution
verified fuzzy
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Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.

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Observation 5e01a35b-472f-414e-8c9a-eb3750893054 · outbound

This paper cites A foundation model for clinical-grade computational pathol- ogy and rare cancers detection.Nature medicine, pages 1–12,.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images A foundation model for clinical-grade computational pathol- ogy and rare cancers detection.Nature medicine, pages 1–12,

Reference 37

Resolution
verified fuzzy
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Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.

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Observation b60b2088-5785-4025-b502-887d6a0ef864 · outbound

This paper cites Allen, Matthew A.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Allen, Matthew A

Reference 38

Resolution
verified fuzzy
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Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.

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Observation 1b3d816d-e40a-4c89-ad89-d261807bcab2 · outbound

This paper cites Spatially re- solved gene expression prediction from histology images via bi-modal contrastive learning.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Spatially re- solved gene expression prediction from histology images via bi-modal contrastive learning

Reference 39

Resolution
verified fuzzy
raw_fallback, observed 2026-08-11T23:24:41.711851Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.

source=pdf_text observed=2026-08-11T23:24:41.506739Z digest=sha256:d43832959d881278dd35edd0521593881a8ad1c4f6d1fa7ca9452872f45dfcbb

Observation 9d1158c8-2e11-4653-8eda-2cf8705a7148 · outbound

This paper cites Exemplar guided deep neural network for spatial tran- scriptomics analysis of gene expression prediction.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Exemplar guided deep neural network for spatial tran- scriptomics analysis of gene expression prediction

Reference 40

Resolution
verified fuzzy
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Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.

source=pdf_text observed=2026-08-11T23:24:41.511818Z digest=sha256:a665a95b606f773fd001b2d501d3857bb2e0c1cabcbe8a1ddab648e3d14a0c11

Observation ee4d8abd-a948-47b7-a477-939406b3a798 · outbound

This paper cites Spatial tran- scriptomics prediction from histology jointly through Trans- former and graph neural networks.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Spatial tran- scriptomics prediction from histology jointly through Trans- former and graph neural networks

Reference 41

Resolution
verified fuzzy
raw_fallback, observed 2026-08-11T23:24:41.677287Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.

source=pdf_text observed=2026-08-11T23:24:41.516482Z digest=sha256:55efeb7121db0680f2214ba2f737de47c9810b1c941aec31def697198249462a

Observation 25152464-3205-477a-b004-b9d9c02414f8 · outbound

This paper cites Experimental Setup The ResNet18-based patch encoder is implemented in Py- Torch (version 2.2.2).

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Experimental Setup The ResNet18-based patch encoder is implemented in Py- Torch (version 2.2.2)

Reference 42

Resolution
verified fuzzy
raw_fallback, observed 2026-08-11T23:24:41.658434Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.

source=pdf_text observed=2026-08-11T23:24:41.521687Z digest=sha256:963e51a4a3f0f118329b67afc7dd70a369a302230b5b68625461c1c849e48e70

Observation ba4be4a4-bf0c-4cee-9207-5829631c1696 · outbound

This paper cites Results Comparison - PCC This section discusses the PCC scores attained by MERGE and TRIPLEX across the samples in ST-Net dataset for two cancer-relevant genes - FASN and GNAS.

MERGE: Multi-faceted Hierarchical Graph-based GNN for Gene Expression Prediction from Whole Slide Histopathology Images Results Comparison - PCC This section discusses the PCC scores attained by MERGE and TRIPLEX across the samples in ST-Net dataset for two cancer-relevant genes - FASN and GNAS

Reference 43

Resolution
verified fuzzy
raw_fallback, observed 2026-08-11T23:24:41.640375Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-17T06:30:58.91139+00:00.

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Pith citing papers

No inbound Pith citation observations are available.