Paper Citation Record · LEDGER

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning

As of 10 August 2026, this Paper Citation Record lists 54 of 54 outbound references and 1 inbound Pith citation observation for arXiv:2502.03499.

A citation records a reference. It does not transfer a finding from one paper to another.

pith.paper-citation-record.v1
2502.03499 v1

Coverage vector

measured 54 of 54 reference resolution

Typed states for the displayed outbound observations.

Source: paper_references, paper_reference_links, observed 2026-08-09T10:16:49.318657Z

measured 55 of 55 standing notices

One-hop event checks from named stored sources.

Source: scholarly_work_events, retraction_status_cache, observed 2026-08-09T06:31:02.800959+00:00

measured 1 of 1 inbound itemization

Pith citing papers itemized under the disclosed page cap.

Source: paper_references, paper_reference_links, observed 2026-08-08T15:15:00.008625Z

measured 0 of 1 external citation measurements

A source-named dated measurement, never combined with another source.

Source: pith, observed 2026-08-08T15:15:00.367295Z

Reference resolution

54 of 54 outbound references displayed

verified exact2
verified fuzzy19
unresolved33
parse uncertain0
malformed identifier0
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External citation measurements

No source-named external measurement is stored.

Outbound references

Observation b8fcacb2-c4f2-44a1-bb66-dde9851378c7 · outbound

This paper cites J., Bambrick, J., et al.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning J., Bambrick, J., et al

Reference 1

Resolution: unresolved; no resolver link, observed 2026-08-09T10:16:49.034944Z

Source-reported events for the cited work

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Observation 6d523a76-f1a3-442b-a3fe-9b0aa599bf87 · outbound

This paper cites T., and Frey, B.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning T., and Frey, B

Reference 2

Resolution: verified fuzzy; raw_fallback, observed 2026-08-09T10:16:50.580343Z

Source-reported events for the cited work

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Observation 82d29707-60b1-4fc0-baaf-d0bc76c4af44 · outbound

This paper cites E., Phalke, N., , Z.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning E., Phalke, N., , Z

Reference 3

Resolution: verified fuzzy; raw_fallback, observed 2026-08-09T10:16:50.563050Z

Source-reported events for the cited work

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Observation 2a62e8e1-a6bb-4e4d-8996-e34a209bc67a · outbound

This paper cites R., Grabska-Barwinska, A., Taylor, K.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning R., Grabska-Barwinska, A., Taylor, K

Reference 4

Resolution: verified fuzzy; raw_fallback, observed 2026-08-09T10:16:50.508368Z

Source-reported events for the cited work

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Observation 48be77e4-5803-4b4e-919b-48685f993478 · outbound

This paper cites Base-resolution models of transcription-factor binding reveal soft motif syntax.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Base-resolution models of transcription-factor binding reveal soft motif syntax

Reference 5

Resolution: verified fuzzy; raw_fallback, observed 2026-08-09T10:16:50.411857Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

Observation ea45ed71-380c-440f-a3a6-8c656699affa · outbound

This paper cites Layer Normalization.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Layer Normalization

Reference 6

Resolution: unresolved; no resolver link, observed 2026-08-09T10:16:49.062404Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

Observation 097a0ac6-0067-484a-b662-604b879ca535 · outbound

This paper cites S., and Song, Y.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning S., and Song, Y

Reference 7

Resolution: unresolved; no resolver link, observed 2026-08-09T10:16:49.069331Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

Observation 4de10e5d-29ac-4961-abae-af40a2131618 · outbound

This paper cites H., Zielinski, M., Sargeant, T., et al.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning H., Zielinski, M., Sargeant, T., et al

Reference 8

Resolution: verified fuzzy; raw_fallback, observed 2026-08-09T10:16:50.283833Z

Source-reported events for the cited work

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Observation 4fe1abeb-0c97-4274-a4ea-fb3132c19753 · outbound

This paper cites Dnalongbench: A benchmark suite for long-range dna prediction tasks.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Dnalongbench: A benchmark suite for long-range dna prediction tasks

Reference 9

Resolution: verified fuzzy; raw_fallback, observed 2026-08-09T10:16:50.253583Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

Observation b12542d4-4c27-4c36-8e50-9c9454dcd949 · outbound

This paper cites H., Oteri, F., Dallago, C., Trop, E., de Almeida, B.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning H., Oteri, F., Dallago, C., Trop, E., de Almeida, B

Reference 10

Resolution: unresolved; no resolver link, observed 2026-08-09T10:16:49.085678Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

Observation e83e057f-1d5d-43da-b545-136c180781a5 · outbound

This paper cites P., Dalla-Torre, H., Richard, G., Blum, C., Hexemer, L., G \'e lard, M., Mendoza-Revilla, J., Pandey, P., Laurent, S., Lopez, M., et al.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning P., Dalla-Torre, H., Richard, G., Blum, C., Hexemer, L., G \'e lard, M., Mendoza-Revilla, J., Pandey, P., Laurent, S., Lopez, M., et al

Reference 11

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Source-reported events for the cited work

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Observation dd3ced72-e136-449a-aecc-705046387110 · outbound

This paper cites Genomic benchmarks: a collection of datasets for genomic sequence classification.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Genomic benchmarks: a collection of datasets for genomic sequence classification

Reference 12

Resolution: verified fuzzy; raw_fallback, observed 2026-08-09T10:16:50.201680Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

Observation 87cd24d4-9bd0-4545-96ad-49b52c043c2d · outbound

This paper cites OLMo: Accelerating the Science of Language Models.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning OLMo: Accelerating the Science of Language Models

Reference 13

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Source-reported events for the cited work

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Observation 05ac99ab-a684-4391-9b93-a9c83877cd64 · outbound

This paper cites Mamba: Linear-Time Sequence Modeling with Selective State Spaces.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Mamba: Linear-Time Sequence Modeling with Selective State Spaces

Reference 14

Resolution: unresolved; no resolver link, observed 2026-08-09T10:16:49.105714Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

Observation 22170b4d-ad7e-48d3-a5a4-631345b1ede8 · outbound

This paper cites Initializing new word embeddings for pretrained language models.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Initializing new word embeddings for pretrained language models

Reference 15

Resolution: verified fuzzy; raw_fallback, observed 2026-08-09T10:16:50.177908Z

Source-reported events for the cited work

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Observation ace1cefb-b4bd-47cc-b07d-466b9b86cd73 · outbound

This paper cites NEFTune: Noisy Embeddings Improve Instruction Finetuning.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning NEFTune: Noisy Embeddings Improve Instruction Finetuning

Reference 16

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Source-reported events for the cited work

Unavailable: canonical work link unavailable.

Observation c5c4e419-1d76-47a1-bfa7-23abb78becbc · outbound

This paper cites Tulu 3: Pushing Frontiers in Open Language Model Post-Training.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Tulu 3: Pushing Frontiers in Open Language Model Post-Training

Reference 17

Resolution: unresolved; no resolver link, observed 2026-08-09T10:16:49.121257Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

Observation ce20b1ce-8537-4d98-8c7e-6c1ddd5f8764 · outbound

This paper cites Genomic data resources: challenges and promises.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Genomic data resources: challenges and promises

Reference 18

Resolution: verified fuzzy; raw_fallback, observed 2026-08-09T10:16:50.159009Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

Observation 5ef8483d-79bd-49f6-85bb-d3239a6afda2 · outbound

This paper cites VQDNA: Unleashing the Power of Vector Quantization for Multi-Species Genomic Sequence Modeling.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning VQDNA: Unleashing the Power of Vector Quantization for Multi-Species Genomic Sequence Modeling

Reference 19

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Source-reported events for the cited work

Unavailable: canonical work link unavailable.

Observation 5c329ddf-d359-46ce-8b42-7f346d4ff5d5 · outbound

This paper cites GV-Rep: A Large-Scale Dataset for Genetic Variant Representation Learning.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning GV-Rep: A Large-Scale Dataset for Genetic Variant Representation Learning

Reference 20

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Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

Observation 533f705a-3b3f-4ca8-a913-26bc8492c904 · outbound

This paper cites an unresolved cited work.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Unresolved cited work

Reference 21

Resolution: unresolved; raw_fallback, observed 2026-08-09T10:16:50.138582Z

Source-reported events for the cited work

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Observation 2df14770-cde3-45f7-9f64-f28dd80ac18c · outbound

This paper cites Reframing Instructional Prompts to GPTk's Language.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Reframing Instructional Prompts to GPTk's Language

Reference 22

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Observation 9fc2c2e4-efc7-41a6-8252-d7faf9a77ed8 · outbound

This paper cites Hyenadna: Long-range genomic sequence modeling at single nucleotide resolution.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Hyenadna: Long-range genomic sequence modeling at single nucleotide resolution

Reference 23

Resolution: unresolved; no resolver link, observed 2026-08-09T10:16:49.153754Z

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Observation 642980e1-6438-4b74-b19e-e13582518198 · outbound

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Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Unresolved cited work

Reference 24

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Source-reported events for the cited work

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Observation 5bb9f469-f237-4676-8224-e6cc4e59f5f3 · outbound

This paper cites Byte Latent Transformer: Patches Scale Better Than Tokens.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Byte Latent Transformer: Patches Scale Better Than Tokens

Reference 25

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Observation f69b1aaa-60d7-44f5-bbb0-cc59be94282c · outbound

This paper cites K., Harbison, C.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning K., Harbison, C

Reference 26

Resolution: verified fuzzy; raw_fallback, observed 2026-08-09T10:16:50.080177Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

Observation 34cd2b42-bb27-45c1-b5c7-a27d16d3fbf6 · outbound

This paper cites Train Short, Test Long: Attention with Linear Biases Enables Input Length Extrapolation.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Train Short, Test Long: Attention with Linear Biases Enables Input Length Extrapolation

Reference 27

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Observation 54c58040-082d-4fc1-bfc6-72ae83cfd73b · outbound

This paper cites Model Decides How to Tokenize: Adaptive DNA Sequence Tokenization with MxDNA.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Model Decides How to Tokenize: Adaptive DNA Sequence Tokenization with MxDNA

Reference 28

Resolution: unresolved; no resolver link, observed 2026-08-09T10:16:49.178190Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

Observation 3e427c02-be63-4eb2-9df0-94c0d2bb067a · outbound

This paper cites Language models are unsupervised multitask learners.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Language models are unsupervised multitask learners

Reference 29

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Observation a695b5cb-d046-43fe-99ba-965e28104a3b · outbound

This paper cites Scaling Language Models: Methods, Analysis & Insights from Training Gopher.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Scaling Language Models: Methods, Analysis & Insights from Training Gopher

Reference 30

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Observation be538b8e-a7f6-46ae-aad5-f28d1001361c · outbound

This paper cites M., and Poetsch, A.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning M., and Poetsch, A

Reference 31

Resolution: verified fuzzy; raw_fallback, observed 2026-08-09T10:16:50.040850Z

Source-reported events for the cited work

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Observation 05fd5130-8003-42f6-a4e9-7dc4f668bef8 · outbound

This paper cites Multitask Prompted Training Enables Zero-Shot Task Generalization.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Multitask Prompted Training Enables Zero-Shot Task Generalization

Reference 32

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Observation 43e134ad-c9db-4374-9ac6-dbdf8713ed31 · outbound

This paper cites Caduceus: Bi-Directional Equivariant Long-Range DNA Sequence Modeling.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Caduceus: Bi-Directional Equivariant Long-Range DNA Sequence Modeling

Reference 33

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Observation 46c99a2f-8ed9-4f45-87ba-611581022801 · outbound

This paper cites Bio-xLSTM: Generative modeling, representation and in-context learning of biological and chemical sequences.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Bio-xLSTM: Generative modeling, representation and in-context learning of biological and chemical sequences

Reference 34

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Observation a6022743-fb3d-45e5-99f5-390ccb298b32 · outbound

This paper cites L., Ciufo, S., Domrachev, M., Hotton, C.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning L., Ciufo, S., Domrachev, M., Hotton, C

Reference 35

Resolution: verified fuzzy; raw_fallback, observed 2026-08-09T10:16:50.018744Z

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Observation bde19eb6-6959-4127-bd5d-91238cd3307c · outbound

This paper cites Neural Machine Translation of Rare Words with Subword Units.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Neural Machine Translation of Rare Words with Subword Units

Reference 36

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Observation 06dee27c-dac4-4f13-92d2-701de7cde316 · outbound

This paper cites Multi-Task Inference: Can Large Language Models Follow Multiple Instructions at Once?.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Multi-Task Inference: Can Large Language Models Follow Multiple Instructions at Once?

Reference 37

Resolution: verified exact; local_arxiv, observed 2026-08-09T10:16:49.465387Z

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Observation 0b7a787a-1e6a-48bd-9d05-89d9d05d9c52 · outbound

This paper cites Roformer: Enhanced transformer with rotary position embedding.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Roformer: Enhanced transformer with rotary position embedding

Reference 38

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Observation cf3d8ac8-918e-49ff-978a-cc2471e115ce · outbound

This paper cites Gemini: A Family of Highly Capable Multimodal Models.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Gemini: A Family of Highly Capable Multimodal Models

Reference 39

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Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning L., Church, G

Reference 40

Resolution: verified fuzzy; raw_fallback, observed 2026-08-09T10:16:49.975206Z

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Observation 1f04c64b-b8c7-4b3a-bc5a-60e764a38e85 · outbound

This paper cites LLaMA: Open and Efficient Foundation Language Models.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning LLaMA: Open and Efficient Foundation Language Models

Reference 41

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Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Neural discrete representation learning

Reference 42

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Observation 97bec217-9f25-4b5e-9206-69c14bf16f18 · outbound

This paper cites J., and Davey, N.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning J., and Davey, N

Reference 43

Resolution: verified fuzzy; raw_fallback, observed 2026-08-09T10:16:49.941118Z

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Observation bc485f35-19eb-4700-8d11-3e752ce2da2f · outbound

This paper cites Attention is all you need.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Attention is all you need

Reference 44

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Observation ee230446-5d57-416a-a0b5-20ecb15acf46 · outbound

This paper cites Multi-purpose rna language modelling with motif-aware pretraining and type-guided fine-tuning.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Multi-purpose rna language modelling with motif-aware pretraining and type-guided fine-tuning

Reference 45

Resolution: verified fuzzy; raw_fallback, observed 2026-08-09T10:16:49.904200Z

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This paper cites V., Zhou, D., et al.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning V., Zhou, D., et al

Reference 46

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Observation 45372b7d-05d7-4399-9386-7321da163d2c · outbound

This paper cites DeepSeek-Prover: Advancing Theorem Proving in LLMs through Large-Scale Synthetic Data.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning DeepSeek-Prover: Advancing Theorem Proving in LLMs through Large-Scale Synthetic Data

Reference 47

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Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning and Bottou, L

Reference 48

Resolution: verified fuzzy; raw_fallback, observed 2026-08-09T10:16:49.864534Z

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Observation c7455a80-611e-4e1a-95a9-7c90c3d1c7fa · outbound

This paper cites C., Sribour, M., Baek, I., Laurin, M., and Fuchs, E.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning C., Sribour, M., Baek, I., Laurin, M., and Fuchs, E

Reference 49

Resolution: verified fuzzy; raw_fallback, observed 2026-08-09T10:16:49.845290Z

Source-reported events for the cited work

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Observation d8d5559a-bcc6-40d7-ac2a-ea8d00d42a5f · outbound

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Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning and Sennrich, R

Reference 50

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Observation 7cc1441a-29b7-473c-8aad-9d882a14b0bd · outbound

This paper cites Towards Lifelong Learning of Large Language Models: A Survey.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning Towards Lifelong Learning of Large Language Models: A Survey

Reference 51

Resolution: unresolved; no resolver link, observed 2026-08-09T10:16:49.301632Z

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Observation 388d891e-00f7-4554-9ef7-efc9b6b70d43 · outbound

This paper cites and Troyanskaya, O.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning and Troyanskaya, O

Reference 52

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This paper cites DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 53

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This paper cites write newline.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning write newline

Reference 54

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Frozen but Not Always Accessible: A Representation Analysis of Genomic Language Models cites this paper.

Frozen but Not Always Accessible: A Representation Analysis of Genomic Language Models Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning

Reference 47

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