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What is active wetting?

T0 review · 0 major / 3 minor · reviewed 2026-08-03 · deepseek-v4-flash

Pith's one-line read The paper proposes restricting the term “active wetting” to wetting phenomena in active liquids, where chemo-mechanical coupling happens at the level of the microscopic bulk constituents, and placing biofilms and purely diffusive condensate

desk verdict A genuinely useful, openly hedged taxonomy for a term that badly needed one—well worth a serious referee. read the letter →

arxiv 2602.10287 v2 pith:GCXGREEY submitted 2026-02-10 cond-mat.soft

classification cond-mat.soft
keywords activewettingclassificationreactiveliquidschemo-mechanicalcouplingBrownianparticlescellmonolayerstransitions
verification ladder T0 review T1 audit T2 compute T3 formal

The pith

A machine-rendered reading of the paper's core claim, the machinery that carries it, and where it could break.

The reading

This paper proposes a working answer to a question that has spread faster than its definition: what counts as “active wetting”? It sets up a coarse classification of wetting phenomena — equilibrium, relaxational, driven, and reactive — and then argues that the label “active wetting” should be reserved for wetting phenomena involving active liquids, meaning systems whose chemo-mechanical coupling operates at the level of the microscopic bulk constituents (self-propelling particles or active stresses). Under that rule, cell monolayers, cell aggregates, and active Brownian-particle layers count; biofilms and purely diffusive biomolecular condensates should instead be called reactive wetting. The paper stresses that the boundary between categories is not fixed but shifts with the level of description chosen.

What carries the argument

The key object is a four-category taxonomy of wetting — equilibrium, relaxational, driven, reactive — used as the backdrop, plus the single discriminating criterion for active wetting: the location of chemo-mechanical coupling (“on the level of the microscopic bulk constituents”). The argument works by surveying the experimental and theoretical systems for which the term “active wetting” is used, showing they split along this line, and then admitting that the line depends on the level of description. The taxonomy itself is the device that makes the definition stateable.

What would settle it

A concrete observation that would undermine the definition: identify a system currently called active wetting (for instance a spreading cell monolayer) whose behaviour is fully reproduced by a model with no bulk-level active stresses — only boundary reactions and passive bulk rheology. If such a system nonetheless shows the characteristic active wetting transitions (e.g., a substrate-stiffness-dependent wetting transition with an intrinsic length scale), then the microscopic-bulk-coupling criterion is not doing the classificatory work claimed.

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Extended reading notes

Core claim

The paper's central claim is a tentative definition: it “could be a clarifying restriction to use the term active wetting only for wetting phenomena involving active liquids, i.e., where the chemo-mechanical coupling takes place on the level of the microscopic bulk constituents.” From this it follows that the category includes the (de)wetting of cell monolayers and aggregates, dense layers of active Brownian particles, and sessile drops of active liquids, while excluding proliferating biofilms and biomolecular condensates described by purely diffusive transport, which fall under reactive wetting. The definition is offered with explicit caveats: the same physical system may appear as active w

Load-bearing premise

The load-bearing premise is that there is a stable, meaningful distinction between chemo-mechanical coupling at the level of the microscopic bulk constituents and couplings that act at other levels — even though the paper itself shows that a single system can be classified either way depending on whether it is described microscopically or via a coarse-grained nonreciprocal field theory.

Editorial extensions

If this is right

  • If the proposed definition catches on, the wetting of biofilms and of purely diffusive condensates will be re-labelled as reactive wetting, removing a current source of ambiguity.
  • The classification gives researchers a shared vocabulary that should make it easier to map contact-angle laws and wetting-transition results across active, reactive, driven, and equilibrium cases.
  • It exposes “active equilibrium wetting” as a contradictory phrase, since active systems are permanently out of equilibrium; that phrase should be replaced by a description of stationary non-equilibrium states.
  • The caveat about description level implies that any experimental claim of “active wetting” should specify whether the microscopic constituents are indeed self-propelling or stress-generating, rather than just out of equilibrium.
  • The list of phenomena surveyed suggests that systematic comparative studies across systems will be needed to extract general laws of dynamic active wetting.

Reading between the lines

Editorial extensions of the paper, not claims the author makes directly.

  • A natural test: take a system that can be modelled both as active Brownian particles and as a nonreciprocal continuum field theory, compute the wetting layer thickness or contact angle in both, and see whether the difference is quantitative or qualitative; if qualitative, the active/reactive distinction may be more than a modelling choice.
  • The criterion might be sharpened by asking whether the chemo-mechanical coupling survives a coarse-graining that keeps the same hydrodynamic variables; if it disappears, the system is arguably reactive at the macroscale.
  • One could imagine an experimental protocol: measure the strength of interface currents (stationary density fluxes) in a wetting layer; if currents are driven by bulk constituent motility rather than by boundary reactions, the phenomenon would be active by the proposed definition.
  • The paper's own caveat hints that the taxonomy is a pragmatic tool, not an ontology; a future synthesis might replace the binary active/reactive label with a parameter describing the scale at which energy injection couples to mechanics.
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Editorial analysis

A structured set of objections, weighed in public.

Desk editor's note, referee report, and a circularity audit.

Referee Report

0 major / 3 minor

Summary. This perspective proposes a coarse taxonomy of wetting phenomena as a way to give a tentative definition of the term "active wetting," which has been used increasingly in the literature without a clear meaning. The author first distinguishes four categories for passive liquids: equilibrium wetting (Section II), relaxational wetting (Section III), driven wetting (Section IV), and reactive wetting (Section V). Section VI then surveys a broad range of recent works that call various phenomena "active wetting," including cell aggregates, epithelial monolayers, biofilms, biomolecular condensates, active Brownian particles, and active liquid drops. The central proposal, presented in Section VII, is to reserve "active wetting" for wetting phenomena involving active liquids, i.e., cases in which chemo-mechanical coupling takes place on the level of the microscopic bulk constituents; under this restriction, biofilms and purely diffusive condensates would fall instead under reactive wetting. The author explicitly stresses that any such classification depends on conceptual idealizations and the chosen level of description, and that the boundary between active and reactive wetting is not intrinsic to the system.

Significance. The paper addresses a real terminological problem in a rapidly growing interdisciplinary field. Its contribution is conceptual rather than quantitative: it offers a coherent framework, a detailed literature mapping, and a specific, falsifiable proposal for what should and should not be called active wetting. The main strength is the explicitly hedged framing: the proposed definition is presented as a clarifying restriction with named caveats, not as a natural-kind statement. The author openly acknowledges the level-of-description ambiguity (Section VII), which is the most serious objection one could raise, and makes a practical recommendation to always clarify the level of description. If accepted, the taxonomy would help standardize discussions across soft-matter physics, biophysics, and active-matter research. The absence of overreach is commendable: the paper does not claim to settle the issue, but to provide a usable starting point. The breadth of references and the careful delineation of overlaps and edge cases make this a useful reference for the community.

minor comments (3)
  1. [Section VII] The proposed definition hinges on the phrase "chemo-mechanical coupling takes place on the level of the microscopic bulk constituents," but "microscopic" is not operationalized. For example, cell aggregates consist of cells that are mesoscopic, not microscopic. Consider clarifying that "microscopic" means "on the scale of the active constituents" (whether molecules, particles, or cells) and explicitly recommend a preferred level of description when using the taxonomy. This would sharpen the central proposal without changing its tentative character.
  2. [Section III, IV, VI, VII] Typos: "chanels" in Section III, "V oinov" in Section IV, "controll parameters" in Section VI, and "mayor difficulties" in Section VII should be corrected to channels, Voinov, control parameters, and major difficulties.
  3. [References] References [107] and [123] appear to be the same article (Morris and Yap, "Wetting by living tissues") with different years (2019 vs. 2018). Please verify and correct the duplicate or the years.

Circularity Check

0 steps flagged · score 0.0 of 10

No significant circularity: the central claim is an explicitly tentative definitional proposal, not a derived prediction.

full rationale

The paper is a perspective that proposes a classification of wetting phenomena and a tentative definition of active wetting. There are no fitted parameters, no equations, and no prediction that is statistically or constructively forced by an input. The central proposal in Section VII — restricting 'active wetting' to wetting by active liquids with chemo-mechanical coupling at the level of microscopic bulk constituents — is presented as a suggestion ('it could be a clarifying restriction') and is explicitly hedged. The author acknowledges that the active/reactive boundary is level-dependent, noting that 'a system could show active wetting when described microscopically via active Brownian particles and run-and-tumble particles and reactive wetting when described macroscopically via a coarse-grained nonreciprocal field theory.' This is not a hidden circular step; it is an open limitation. Self-citations appear throughout, but they are used as illustrative examples of prior modeling work rather than as load-bearing proof of the proposed definition. No uniqueness theorem or ansatz is imported from the author's own prior work to force the conclusion. The paper's contribution is a stipulative taxonomy, and its caveats prevent the definition from being circular.

Assumptions & free parameters 0 free parameters · 3 assumptions · 0 invented entities

The central proposal rests on the prior notion of 'active liquids' and on the assumption that a coarse taxonomy is useful even though the paper acknowledges its boundaries depend on idealizations. No free parameters or invented entities.

assumptions (3)
  • domain assumption Active liquid is defined as liquid consisting of self-propelled constituents and/or featuring active stresses resulting from properties of the constituents.
    The proposed definition of active wetting relies on this prior notion, citing Refs. [16-19], but the exact boundary of 'active liquid' is not formalized.
  • domain assumption The classification into equilibrium, relaxational, driven, and reactive wetting is meaningful and covers the relevant phenomena.
    The paper accepts that boundaries depend on idealizations and can be ambiguous (Section VII), yet still uses them as the basis for the definition.
  • ad hoc to paper The level of description (microscopic bulk constituents vs. coarse-grained field) is a stable criterion to decide 'active' vs 'reactive' wetting.
    The author explicitly proposes this criterion and immediately notes in Section VII that the same system can be classified differently at different description levels.

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Cite this review

Pith. "Pith review of What is active wetting?." pith.science (2026). https://pith.science/paper/GCXGREEY

@misc{pith2026260210287,
  author       = {Pith},
  title        = {Pith review of: What is active wetting?},
  year         = {2026},
  howpublished = {\url{https://pith.science/paper/GCXGREEY}},
  note         = {Machine review of arXiv:2602.10287}
}
read the original abstract

In recent years the term \textit{active wetting} has gained some traction in works describing, analyzing and modeling a wide variety of wetting phenomena, for instance, in the contexts of biomolecular condensates, of cell layers or cell aggregates, and of active Brownian particles. The present perspective discusses a coarse classification of wetting phenomena that accounts for this. First, different categories of static and dynamic wetting of passive liquids are briefly introduced, in particular, distinguishing equilibrium wetting, relaxational wetting, driven wetting, and reactive wetting. Second, an overview is given of the various phenomena recently described as active wetting. We conclude by discussing a possible definition of active wetting together with a number of caveats that one might want to keep in mind when using such classifications.

Figures

Figures reproduced from arXiv: 2602.10287 by the authors.

Figure 1
Figure 1. FIG. 1: Sketches of equilibrium wetting: (a) complete wetting, (b) [PITH_FULL_IMAGE:figures/full_fig_p001_1.png] view at source ↗
Figure 3
Figure 3. FIG. 3: Sketches of driven wetting: (a) drop sliding down an incline, [PITH_FULL_IMAGE:figures/full_fig_p002_3.png] view at source ↗
Figure 2
Figure 2. FIG. 2: Sketches of relaxational wetting: (a) spreading drop (b) [PITH_FULL_IMAGE:figures/full_fig_p002_2.png] view at source ↗
Figures from the paper (2 more)
Figure 4
Figure 4. Figure 4: FIG. 4: Sketches of reactive wetting: (a) moving drop self-propelled [PITH_FULL_IMAGE:figures/full_fig_p003_4.png]
Figure 5
Figure 5. Figure 5: FIG. 5: Sketches of active wetting: (a) static active wetting layer at [PITH_FULL_IMAGE:figures/full_fig_p004_5.png]

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Forward citations

Cited by 1 Pith paper

Reviewed papers in the Pith corpus that reference this work. Sorted by Pith novelty score. Full citation record

  1. Interface-dominated sliding compound drops

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    Stationary sliding compound drops exist only in finite parameter ranges; the 2-1 configuration always slides faster than the 1-2 configuration, and beyond saddle-node bifurcations they undergo periodic fusion-overtaki...

Reference graph

Works this paper leans on

208 extracted references · 105 canonical work pages · cited by 1 Pith paper

  1. [1]

    W. B. Hardy. Historical notes upon surface energy and forces of short range.Nature, 109:375–378, 1922. doi: 10.1038/ 109375a0

  2. [3]

    Hauksbee

    F. Hauksbee. An account of an experiment touching the pro- portions of the ascent of spirit of wine between two glass planes, whose surfaces were plac’d at certain different dis- tances from each other.Phil. Trans., 28:151–152, 1713. doi: 0.1098/rstl.1713.0012

  3. [4]

    J. A. Segner. De figuris superficierum fluidarum.Comm. Soc. Reg. Sci. Gottingensis, 1:301, 1751

  4. [5]

    T. Young. An essay on the cohesion of fluids.Phil. Trans. R. Soc., 95:65–87, 1805. doi: 10.1098/rstl.1805.0005

  5. [6]

    P. S. Laplace. Supplement of volume X: Sur l’action capillaire. InTrait ´e de M´ecanique C´eleste. 1806

  6. [7]

    Newton.Opticks

    I. Newton.Opticks. G. Bell & Sons LTD., London, 1730. Preprint– contact: u.thiele@uni-muenster.de – www.uwethiele.de – February 12, 2026 8 (reprinted 4th ed. 1931)

  7. [8]

    Rowlinson and B

    J. Rowlinson and B. Widom.Molecular theory of capillarity. Oxford University Press, Oxford, 1982

  8. [9]

    P. G. de Gennes. Wetting: Statics and dynamics.Rev. Mod. Phys., 57:827–863, 1985. doi: 10.1103/RevModPhys.57.827

Show all 208 references
  1. [10]

    de Gennes, F

    P.-G. de Gennes, F. Brochard-Wyart, and D. Qu´er´e.Capillar- ity and Wetting Phenomena: Drops, Bubbles, Pearls, Waves. Springer, New York, 2004. ISBN 978-0-387-21656-0. doi: 10.1007/978-0-387-21656-0

  2. [11]

    V . M. Starov, M. G. Velarde, and C. J. Radke.Wetting and spreading dynamics. Taylor and Francis, Boca Raton, 2007

  3. [12]

    D. Bonn, J. Eggers, J. Indekeu, J. Meunier, and E. Rolley. Wet- ting and spreading.Rev. Mod. Phys., 81:739–805, 2009. doi: 10.1103/RevModPhys.81.739

  4. [13]

    R. V . Craster and O. K. Matar. Dynamics and stability of thin liquid films.Rev. Mod. Phys., 81:1131–1198, 2009. doi: 10. 1103/RevModPhys.81.1131

  5. [14]

    Academic Press (imprint of Elsevier), Amster- dam, 2015

    David Brutin.Droplet wetting and evaporation : from pure to complex fluids. Academic Press (imprint of Elsevier), Amster- dam, 2015. ISBN 9780128007228

  6. [15]

    E. Y . Bormashenko.Physics of wetting: phenomena and ap- plications of fluids on surfaces. De Gruyter, Berlin/Boston,

  7. [16]

    M. C. Marchetti, J. F. Joanny, S. Ramaswamy, T. B. Liverpool, J. Prost, M. Rao, and R. A. Simha. Hydrodynamics of soft active matter.Rev. Mod. Phys., 85:1143–1189, 2013. doi: 10.1103/RevModPhys.85.1143

  8. [17]

    S. R. Nagel. Experimental soft-matter science.Rev. Mod. Phys., 89:025002, 2017. doi: 10.1103/revmodphys.89. 025002

  9. [18]

    M. J. Bowick, N. Fakhri, M. C. Marchetti, and S. Ramaswamy. Symmetry, thermodynamics, and topology in active matter. Phys. Rev. X, 12:010501, 2022. doi: 10.1103/PhysRevX.12. 010501

  10. [20]

    Hinrichsen

    H. Hinrichsen. Non-equilibrium phase transitions.Physica A, 369:1–28, 2006. doi: 10.1016/j.physa.2006.04.007

  11. [21]

    Dietrich

    S. Dietrich. Wetting phenomena. In C. Domb and J. L. Lebowitz, editors,Phase Transitions and Critical Phenomena, volume 12, pages 1–218. Academic Press, London, 1988

  12. [22]

    J. O. Indekeu. Wetting phase transitions and critical phenom- ena in condensed matter.Physica A, 389:4332–4359, 2010. doi: 10.1016/j.physa.2010.02.017

  13. [23]

    A. O. Parry, C. Rascon, E. A. G. Jamie, and D. G. A. L. Aarts. Capillary emptying and short-range wetting.Phys. Rev. Lett., 108:246101, 2012. doi: 10.1103/PhysRevLett.108.246101

  14. [24]

    Qu ´er´e

    D. Qu ´er´e. Wetting and roughness.Ann. Rev. Mater. Res., 38:71–99, 2008. doi: 10.1146/annurev.matsci.38.060407. 132434

  15. [25]

    Andreotti and J

    B. Andreotti and J. H. Snoeijer. Statics and dynamics of soft wetting.Annu. Rev. Fluid Mech., 52:285–308, 2020. doi: 10.1146/annurev-fluid-010719-060147

  16. [27]

    Woywod and M

    D. Woywod and M. Schoen. The wetting of planar solid sur- faces by symmetric binary mixtures near bulk gas-liquid coex- istence.J. Phys.: Condens. Matter, 16:4761–4783, 2004. doi: 10.1088/0953-8984/16/28/002

  17. [28]

    Fondecave and F

    R. Fondecave and F. Brochard-Wyart. Wetting laws for poly- mer solutions.Europhys. Lett., 37:115–120, 1997. doi: 10.1209/epl/i1997-00120-y

  18. [29]

    L. H. Tanner. The spreading of silicone oil drops on hor- izontal surfaces.J. Phys. D, 12:1473–1484, 1979. doi: 10.1088/0022-3727/12/9/009

  19. [30]

    H.-J. Butt, J. Liu, K. Koynov, C. Straub, B.and Hinduja, I. Roismann, R. Berger, X. Li, D. V ollmer, W. Steffen, and M. Kappl. Contact angle hysteresis.Curr. Opin. Colloid Inter- face Sci., 59:101574, 2022. doi: 10.1016/j.cocis.2022.101574

  20. [32]

    Kondic, A

    L. Kondic, A. G. Gonz ´alez, J. A. Diez, J. D. Fowlkes, and P. Rack. Liquid-state dewetting of pulsed-laser- heated nanoscale metal films and other geometries.Ann. Rev. Fluid Mech., 52:235–262, 2020. doi: 10.1146/ annurev-fluid-010719-060340

  21. [33]

    T. R. Kotni, J. Sarkar, and R. Khanna. Dewetting of thin wetting film supported by different solid substrates: a review. Phase Transit., 95:551–566, 2022. doi: 10.1080/01411594. 2022.2094267

  22. [34]

    Sch ¨affer and P

    E. Sch ¨affer and P. Z. Wong. Contact line dynamics near the pinning threshold: A capillary rise and fall experiment.Phys. Rev. E, 61:5257–5277, 2000. doi: 10.1103/physreve.61.5257

  23. [35]

    J. Bico, E. Reyssat, and B. Roman. Elastocapillar- ity: When surface tension deforms elastic solids.Annu. Rev. Fluid Mech., 50:629–659, 2018. doi: 10.1146/ annurev-fluid-122316-050130

  24. [36]

    H.-J. Butt, R. Berger, W. Steffen, D. V ollmer, and S. A. L. Weber. Adaptive wetting - adaptation in wetting.Langmuir, 34:11292–11304, 2018. doi: 10.1021/acs.langmuir.8b01783

  25. [37]

    L. Q. Chen, E. Bonaccurso, T. Gambaryan-Roisman, V . Starov, N. Koursari, and Y . P. Zhao. Static and dynamic wetting of soft substrates.Curr. Opin. Colloid Interface Sci., 36:46–57, 2018. doi: 10.1016/j.cocis.2017.12.001

  26. [38]

    L. I. S. Mensink, S. de Beer, and J. H. Snoeijer. The role of entropy in wetting of polymer brushes.Soft Matter, 17:1368– 1375, 2021. doi: 10.1039/d0sm00156b

  27. [39]

    S. A. Etha, P. R. Desai, H. S. Sachar, and S. Das. Wetting dynamics on solvophilic, soft, porous, and responsive sur- faces.Macromolecules, 54:584–596, 2021. doi: 10.1021/acs. macromol.0c02234

  28. [40]

    Hartmann, J

    S. Hartmann, J. Diekmann, D. Greve, and U. Thiele. Drops on polymer brushes – Advances in thin-film modelling of adaptive substrates.Langmuir, 40:4001–4021, 2024. doi: 10.1021/acs.langmuir.3c03313

  29. [41]

    Gambaryan-Roisman

    T. Gambaryan-Roisman. Liquids on porous layers: wetting, imbibition and transport processes.Curr. Opin. Colloid In- terface Sci., 19:320–335, 2014. doi: 10.1016/j.cocis.2014.09. 001

  30. [42]

    Johnson, A

    P. Johnson, A. Trybala, and V . Starov. Kinetics of spreading over porous substrates.Colloid Interfac., 3:38, 2019. doi: 10.3390/colloids3010038

  31. [43]

    Hartmann and U

    S. Hartmann and U. Thiele. Gradient dynamics model for drops of volatile liquid on a porous substrate.Phys. Rev. Fluids, 10:014003, 2025. doi: 10.1103/PhysRevFluids.10. 014003

  32. [44]

    M. M. Flapper, A. Pandey, M. H. Essink, E. H. V . Brummelen, S. Karpitschka, and J. H. Snoeijer. Reversal of solvent migra- tion in poroelastic folds.Phys. Rev. Lett., 130:228201, 2023. doi: 10.1103/PhysRevLett.130.228201

  33. [45]

    G. F. Teletzke, H. T. Davis, and L. E. Scriven. Wetting hydro- dynamics.Rev. Phys. Appl. (Paris), 23:989–1007, 1988. doi: 10.1051/rphysap:01988002306098900. Preprint– contact: u.thiele@uni-muenster.de – www.uwethiele.de – February 12, 2026 9

  34. [46]

    J. H. Snoeijer and B. Andreotti. Moving contact lines: Scales, regimes, and dynamical transitions.Annu. Rev. Fluid Mech., 45:269–292, 2013. doi: 10.1146/ annurev-fluid-011212-140734

  35. [47]

    Huh and L

    C. Huh and L. E. Scriven. Hydrodynamic model of steady movement of a solid / liquid / fluid contact line.J. Colloid Interface Sci., 35:85–101, 1971. doi: 10.1016/0021-9797(71) 90188-3

  36. [48]

    Podgorski, J.-M

    T. Podgorski, J.-M. Flesselles, and L. Limat. Corners, cusps, and pearls in running drops.Phys. Rev. Lett., 87:036102, 2001. doi: 10.1103/PhysRevLett.87.036102

  37. [49]

    Engelnkemper, M

    S. Engelnkemper, M. Wilczek, S. V . Gurevich, and U. Thiele. Morphological transitions of sliding drops - dynamics and bi- furcations.Phys. Rev. Fluids, 1:073901, 2016. doi: 10.1103/ PhysRevFluids.1.073901

  38. [50]

    F. Melo, J. F. Joanny, and S. Fauve. Fingering instability of spinning drops.Phys. Rev. Lett., 63:1958–1961, 1989. doi: 10.1103/physrevlett.63.1958

  39. [51]

    J. A. Diez and L. Kondic. Contact line instabilities of thin liquid films.Phys. Rev. Lett., 86:632–635, 2001. doi: 10. 1103/PhysRevLett.86.632

  40. [52]

    A. M. Cazabat, F. Heslot, S. M. Troian, and P. Carles. Fin- gering instability of thin spreading films driven by tempera- ture gradients.Nature, 346:824–826, 1990. doi: 10.1038/ 346824a0

  41. [53]

    M. K. Chaudhury and G. M. Whitesides. How to make wa- ter run uphill.Science, 256:1539–1541, 1992. doi: 10.1126/ science.256.5063.1539

  42. [54]

    Bueno, Y

    J. Bueno, Y . Bazilevs, R. Juanes, and H. Gomez. Droplet motion driven by tensotaxis.Extreme Mech. Lett., 13:10–16,

  43. [55]

    Zhang and T

    Z. Zhang and T. Qian. Variational approach to droplet trans- port via bendotaxis: Thin film dynamics and model reduc- tion.Phys. Rev. Fluids, 7(4):044002, 2022. doi: 10.1103/ physrevfluids.7.044002

  44. [56]

    Barrio-Zhang, ´E Ruiz-Guti ´errez, D

    H. Barrio-Zhang, ´E Ruiz-Guti ´errez, D. Orejon, G. G. Wells, and R. Ledesma-Aguilar. Droplet motion driven by humidity gradients during evaporation and condensation.Eur. Phys. J. E, 47:32, 2024. doi: 10.1140/epje/s10189-024-00426-7

  45. [57]

    Rio and F

    E. Rio and F. Boulogne. Withdrawing a solid from a bath: how much liquid is coated?Adv. Colloid Interfac., 247:100–114, September 2017. doi: 10.1016/j.cis.2017.01.006

  46. [58]

    Tewes, M

    W. Tewes, M. Wilczek, S. V . Gurevich, and U. Thiele. Self- organised dip-coating patterns of simple, partially wetting, nonvolatile liquids.Phys. Rev. Fluids, 4:123903, 2019. doi: 10.1103/PhysRevFluids.4.123903

  47. [59]

    K. B. Blodgett. Films built by depositing successive monomolecular layers on a solid surface.J. Am. Chem. Soc., 57:1007–1022, 1935. doi: 10.1021/ja01309a011

  48. [60]

    M. H. K ¨opf, S. V . Gurevich, R. Friedrich, and U. Thiele. Substrate-mediated pattern formation in monolayer transfer: a reduced model.New J. Phys., 14:023016, 2012. doi: 10.1088/1367-2630/14/2/023016

  49. [61]

    O. N. Oliveira, L. Caseli, and K. Ariga. The past and the future of Langmuir and Langmuir–Blodgett films.Chem. Rev., 122: 6459–6513, 2022. doi: 10.1021/acs.chemrev.1c00754

  50. [62]

    Schubotz, C

    S. Schubotz, C. Honnigfort, S. Nazari, A. Fery, J.-U. Som- mer, P. Uhlmann, B. Braunschweig, and G. K. Auernhammer. Memory effects in polymer brushes showing co-nonsolvency effects.Adv. Colloid Interface Sci., 294:102442, 2021. doi: 10.1016/j.cis.2021.102442

  51. [63]

    K. J. Ruschak. Coating flows.Annu. Rev. Fluid Mech., 17: 65–89, 1985. doi: 10.1146/annurev.fl.17.010185.000433

  52. [64]

    Kajiya, A

    T. Kajiya, A. Daerr, T. Narita, L. Royon, F. Lequeux, and L. Limat. Advancing liquid contact line on visco-elastic gel substrates: Stick-slip vs. continuous motions.Soft Matter, 9: 454–461, 2013. doi: 10.1039/c2sm26714d

  53. [65]

    Mokbel, S

    D. Mokbel, S. Aland, and S. Karpitschka. Stick-slip con- tact line motion on Kelvin-Voigt model substrates.Europhys. Lett., 139:33002, 2022. doi: 10.1209/0295-5075/ac6ca6

  54. [67]

    Le Grand, A

    N. Le Grand, A. Daerr, and L. Limat. Shape and motion of drops sliding down an inclined plane.J. Fluid Mech., 541: 293–315, 2005. doi: 10.1017/s0022112005006105

  55. [68]

    Dreyer and F

    K. Dreyer and F. R. Hickey. The route to chaos in a dripping water faucet.Am. J. Phys., 59:619–627, 1991. doi: 10.1119/ 1.16783

  56. [69]

    Ziegler, J

    J. Ziegler, J. H. Snoeijer, and J. Eggers. Film transitions of receding contact lines.Eur. Phys. J.-Spec. Top., 166:177–180,

  57. [70]

    Galvagno, D

    M. Galvagno, D. Tseluiko, H. Lopez, and U. Thiele. Con- tinuous and discontinuous dynamic unbinding transitions in drawn film flow.Phys. Rev. Lett., 112:137803, 2014. doi: 10.1103/PhysRevLett.112.137803

  58. [71]

    Mohammad Karim

    A. Mohammad Karim. A review of physics of moving con- tact line dynamics models and its applications in interfacial science.J. Appl. Phys., 132, 2022. doi: 10.1063/5.0102028

  59. [72]

    D. Peschka. Variational approach to dynamic contact angles for thin films.Phys. Fluids, 30:082115, 2018. doi: 10.1063/1. 5040985

  60. [73]

    A. V . Lyushnin, A. A. Golovin, and L. M. Pismen. Fingering instability of thin evaporating liquid films.Phys. Rev. E, 65: 021602, 2002. doi: 10.1103/PhysRevE.65.021602

  61. [74]

    Han and Z

    W. Han and Z. Lin. Learning from ”Coffee Rings”: Ordered structures enabled by controlled evaporative self-assembly. Angew. Chem. Int. Ed., 51:1534–1546, 2012. doi: 10.1002/ anie.201104454

  62. [75]

    R. G. Larson. Transport and deposition patterns in dry- ing sessile droplets.Aiche J., 60:1538–1571, 2014. doi: 10.1002/aic.14338

  63. [76]

    U. Thiele. Patterned deposition at moving contact line.Adv. Colloid Interface Sci., 206:399–413, 2014. doi: 10.1016/j.cis. 2013.11.002

  64. [77]

    Lazar and H

    P. Lazar and H. Riegler. Reversible self-propelled droplet movement: a new driving mechanism.Phys. Rev. Lett., 95: 136103, 2005. doi: 10.1103/PhysRevLett.95.136103

  65. [78]

    Yochelis and L

    A. Yochelis and L. M. Pismen. Droplet motion driven by surface freezing or melting: A mesoscopic hydrodynamic ap- proach.Phys. Rev. E, 72:025301(R), 2005. doi: 10.1103/ PhysRevE.72.025301

  66. [79]

    Kumar and K

    G. Kumar and K. N. Prabhu. Review of non-reactive and reac- tive wetting of liquids on surfaces.Adv. Colloid Interface Sci., 133:61–89, 2007. doi: 10.1016/j.cis.2007.04.009

  67. [80]

    Shioi, T

    A. Shioi, T. Ban, and Y . Morimune. Autonomously moving colloidal objects that resemble living matter.Entropy, 12: 2308–2332, 2010. doi: 10.3390/e12112308

  68. [81]

    Eustathopoulos and R

    N. Eustathopoulos and R. V oytovych. The role of reactivity in wetting by liquid metals: A review.J. Mater. Sci., 51:425–437,

  69. [82]

    Domingues Dos Santos and T

    F. Domingues Dos Santos and T. Ondarc ¸uhu. Free-running droplets.Phys. Rev. Lett., 75:2972–2975, 1995. doi: 10.1103/ PhysRevLett.75.2972

  70. [83]

    Brochard-Wyart and P.-G

    F. Brochard-Wyart and P.-G. de Gennes. Spontaneous motion of a reactive droplet.C. R. Acad. Sci. Ser. II, 321:285–288, 1995

  71. [84]

    Sumino, H

    Y . Sumino, H. Kitahata, K. Yoshikawa, M. Nagayama, S. M. Preprint– contact: u.thiele@uni-muenster.de – www.uwethiele.de – February 12, 2026 10 Nomura, N. Magome, and Y . Mori. Chemosensitive run- ning droplet.Phys. Rev. E, 72:041603, 2005. doi: 10.1103/ PhysRevE.72.041603

  72. [86]

    V oss and U

    F. V oss and U. Thiele. Gradient dynamics approach to reactive thin-film hydrodynamics.J. Eng. Math., 149:2/1–40, 2024. doi: 10.1007/s10665-024-10402-x

  73. [87]

    L. E. Scriven and C. V . Sternling. Marangoni effects.Nature, 187:186–188, 1960. doi: 10.1038/187186a0

  74. [88]

    V oss and U

    F. V oss and U. Thiele. Chemomechanical motility modes of partially wetting liquid droplets.Phys. Rev. Fluids, 10:094005,

  75. [89]

    V oss and U

    F. V oss and U. Thiele. From bipedal to chaotic motion of chemically fueled partially wetting liquid drops. 2025. preprint at http://arxiv.org/abs/2512.14370

  76. [90]

    ac- tive wetting

    similar to phenomena described for the case of chemi- cal reactions [89]. Tentatively, one could say that reactive wetting results from imposed persistent gradients that are not spatial along the sub- strate as in section IV but between reservoirs of different prop- erties. Ex...

  77. [91]

    Karapetsas, R

    G. Karapetsas, R. V . Craster, and O. K. Matar. On surfactant- enhanced spreading and superspreading of liquid drops on solid surfaces.J. Fluid Mech., 670:5–37, 2011. doi: 10.1017/ S0022112010005495

  78. [92]

    Thiele and E

    U. Thiele and E. Knobloch. Thin liquid films on a slightly inclined heated plate.Physica D, 190:213–248, 2004. doi: 10.1016/j.physd.2003.09.048

  79. [93]

    Todorova, U

    D. Todorova, U. Thiele, and L. M. Pismen. The relation of steady evaporating drops fed by an influx and freely evapo- rating drops.J. Eng. Math., 73:17–30, 2012. doi: 10.1007/ s10665-011-9485-1

  80. [94]

    Pahlavan and M

    A. Pahlavan and M. Murrell. Active wetting: statics and dy- namics.Annu. Rev. Conden. Ma. P ., 2025. doi: 10.1146/ annurev-conmatphys-061225-105656

  81. [95]

    Pimienta and C

    V . Pimienta and C. Antoine. Self-propulsion on liquid sur- faces.Curr. Opin. Colloid Interface Sci., 19:290–299, 2014. doi: 10.1016/j.cocis.2014.04.001

  82. [96]

    Maitra, P

    A. Maitra, P. Srivastava, M. Rao, and S. Ramaswamy. Acti- vating membranes.Phys. Rev. Lett., 112:258101, 2014. doi: 10.1103/PhysRevLett.112.258101

  83. [97]

    Douezan, K

    S. Douezan, K. Guevorkian, R. Naouar, S. Dufour, D. Cuve- lier, and F. Brochard-Wyart. Spreading dynamics and wetting transition of cellular aggregates.Proc. Natl. Acad. Sci. U. S. A., 108:7315–7320, 2011. doi: 10.1073/pnas.1018057108

  84. [98]

    Douezan, J

    S. Douezan, J. Dumond, and F. Brochard-Wyart. Wetting tran- sitions of cellular aggregates induced by substrate rigidity.Soft Matter, 8:4578–4583, 2012. doi: 10.1039/c2sm07418d

  85. [99]

    Beaune, T

    G. Beaune, T. V . Stirbat, N. Khalifat, O. Cochet-Escartin, S. Garcia, V . V . Gurchenkov, M. P. Murrell, S. Dufour, D. Cu- velier, and F. Brochard-Wyart. How cells flow in the spread- ing of cellular aggregates.Proc. Natl. Acad. Sci. U. S. A., 111: 8055–8060, 2014. doi: 10.10...

  86. [100]

    J. F. Joanny, K. Kruse, J. Prost, and S. Ramaswamy. The actin cortex as an active wetting layer.Eur. Phys. J. E, 36:52, 2013. doi: 10.1140/epje/i2013-13052-9

  87. [101]

    M. H. K ¨opf and L. M. Pismen. A continuum model of ep- ithelial spreading.Soft Matter, 9:3727–3734, 2013. doi: 10.1039/c3sm26955h

  88. [102]

    M. H. K ¨opf and L. M. Pismen. Non-equilibrium patterns in polarizable active layers.Physica D, 259:48–54, 2013. doi: 10.1016/j.physd.2013.05.009

  89. [104]

    Blanch-Mercader and J

    C. Blanch-Mercader and J. Casademunt. Hydrodynamic in- stabilities, waves and turbulence in spreading epithelia.Soft Matter, 13:6913–6928, 2017. doi: 10.1039/c7{s}m01128h

  90. [105]

    Pi-Jauma, R

    I. Pi-Jauma, R. Alert, and J. Casademunt. Collective durotaxis of cohesive cell clusters on a stiffness gradient.Eur. Phys. J. E, 45:7, 2022. doi: 10.1140/epje/s10189-021-00150-6

  91. [106]

    P ´erez-Gonz´alez, R

    C. P ´erez-Gonz´alez, R. Alert, C. Blanch-Mercader, M. G´omez- Gonz´alez, T. Kolodziej, E. Bazellieres, J. Casademunt, and X. Trepat. Active wetting of epithelial tissues.Nat. Phys., 15: 79–88, 2019. doi: 10.1038/s41567-018-0279-5

  92. [108]

    Trenado, L

    C. Trenado, L. L. Bonilla, and A. Mart ´ınez-Calvo. Finger- ing instability in spreading epithelial monolayers: roles of cell polarisation, substrate friction and contractile stresses.Soft Matter, 17:8276–8290, 2021. doi: 10.1039/d1{s}m00626f

  93. [109]

    Morita, S

    H. Morita, S. Grigolon, M. Bock, S. F. G. Krens, G. Salbreux, and C. P. Heisenberg. The physical basis of coordinated tissue spreading in zebrafish gastrulation.Dev. Cell, 40:354–366,

  94. [110]

    Blanch-Mercader, R

    C. Blanch-Mercader, R. Vincent, E. Bazellieres, X. Serra- Picamal, X. Trepat, and J. Casademunt. Effective viscosity and dynamics of spreading epithelia: a solvable model.Soft Matter, 13:1235–1243, 2017. doi: 10.1039/c6sm02188c

  95. [111]

    Lemahieu, P

    G. Lemahieu, P. Moreno-Layseca, T. Hub, C. Bevilacqua, M. G´omez-Gonz´alez, F. Pennarola, F. Colombo, A. E. Massey, L. Barzaghi, A. Palamidessi, L. L. Homagk, S. F. H. Bar- nett, A. X. Cartagena-Rivera, C. Selhuber-Unkel, R. Prevedel, X. Trepat, J. P. Spatz, J. Ivaska, G. Scit...

  96. [112]

    Alert and J

    R. Alert and J. Casademunt. Role of substrate stiffness in tis- sue spreading: wetting transition and tissue durotaxis.Lang- muir, 35:7571–7577, 2019. doi: 10.1021/acs.langmuir.8{b} 02037

  97. [113]

    R. Alert. Fingering instability of active nematic droplets. J. Phys. A-Math. Theor., 55:234009, 2022. doi: 10.1088/ 1751-8121/ac6c61

  98. [115]

    doi: 10.1016/j.devcel.2017.01.010

  99. [116]

    Wallmeyer, S

    B. Wallmeyer, S. Trinschek, S. Yigit, U. Thiele, and T. Betz. Collective cell migration in embryogenesis follows the laws of wetting.Biophys. J., 114:213–222, 2018. doi: 10.1016/j.bpj. 2017.11.011

  100. [117]

    X. Y . Wang, D. Gonzalez-Rodriguez, T. V ourc’h, P. Silberzan, and A. I. Barakat. Contractility-induced self-organization of smooth muscle cells: from multilayer cell sheets to dynamic three-dimensional clusters.Commun. Biol., 6:262, 2023. doi: 10.1038/s42003-023-04578-8

  101. [118]

    M. E. Pallar ´es, I. Pi-Jaum ´a, I. C. Fortunato, V . Grazu, M. G ´omez-Gonz´alez, P. Roca-Cusachs, J. M. De La Fuente, Preprint– contact: u.thiele@uni-muenster.de – www.uwethiele.de – February 12, 2026 11 R. Alert, R. Sunyer, J. Casademunt, and X. Trepat. Stiffness- dependent...

  102. [119]

    H. Z. Ford, G. L. Celora, E. R. Westbrook, M. P. Dalwadi, B. J. Walker, H. Baumann, C. J. Weijer, P. Pearce, and J. R. Chubb. Pattern formation along signaling gradients driven by active droplet behavior of cell swarms.Proc. Natl. Acad. Sci., 122, 2025. doi: 10.1073/pnas.2419152122

  103. [120]

    Reiter and A

    G. Reiter and A. Sharma. Auto-optimization of dewetting rates by rim instabilities in slipping polymer films.Phys. Rev. Lett., 87:166103, 2001. doi: 10.1103/PhysRevLett.87.166103

  104. [121]

    M. E. Black and J. W. Shaevitz. Rheological dynamics of activeMyxococcus xanthuspopulations during develop- ment.Phys. Rev. Lett., 130:218402, 2023. doi: 10.1103/ PhysRevLett.130.218402

  105. [122]

    T. R. Huycke, T. J. H ¨akkinen, H. Miyazaki, V . Srivastava, E. Barruet, C. S. Mcginnis, A. Kalantari, J. Cornwall-Scoones, D. Vaka, Q. Zhu, H. Jo, R. Oria, V . M. Weaver, W. F. De- grado, M. Thomson, K. Garikipati, D. Boffelli, O. D. Klein, and Z. J. Gartner. Patterning and f...

  106. [123]

    R. G. Morris and A. S. Yap. Wetting by living tissues.Nat. Phys., 15:6–7, 2018. doi: 10.1038/s41567-018-0316-4

  107. [124]

    Alert and X

    R. Alert and X. Trepat. Living cells on the move.Phys. Today, 74:30–36, 2021. doi: 10.1063/pt.3.4770

  108. [125]

    Pajic-Lijakovic and M

    I. Pajic-Lijakovic and M. Milivojevic. Active wetting of epithelial tissues: modeling considerations.Eur. Bio- phys. J. Biophys. Lett., 52:1–15, 2023. doi: 10.1007/ s00249-022-01625-w

  109. [126]

    M. A. Fardin, O. M. Rossier, P. Rangamani, P. D. Avigan, N. C. Gauthier, W. V onnegut, A. Mathur, J. Hone, R. Iyengar, and M. P. Sheetz. Cell spreading as a hydrodynamic process. Soft Matter, 6:4788–4799, 2010. doi: 10.1039/c0sm00252f

  110. [127]

    Beaune, C

    G. Beaune, C. Blanch-Mercader, S. Douezan, J. Dumond, D. Gonzalez-Rodriguez, D. Cuvelier, T. Ondarcuhu, P. Sens, S. Dufour, M. P. Murrell, and F. Brochard-Wyart. Sponta- neous migration of cellular aggregates from giant keratocytes to running spheroids.Proc. Natl. Acad. Sci. U...

  111. [128]

    Gonzalez-Rodriguez, K

    D. Gonzalez-Rodriguez, K. Guevorkian, S. Douezan, and F. Brochard-Wyart. Soft matter models of developing tis- sues and tumors.Science, 338:910–917, 2012. doi: 10.1126/ science.1226418

  112. [129]

    Verstraeten, K

    N. Verstraeten, K. Braeken, B. Debkumari, M. Fauvart, J. Fransaer, J. Vermant, and J. Michiels. Living on a surface: swarming and biofilm formation.Trends Microbiol., 16:496– 506, 2008. doi: 10.1016/j.tim.2008.07.004

  113. [130]

    T. E. Angelini, M. Roper, R. Kolter, D. A. Weitz, and M. P. Brenner.Bacillus subtilisspreads by surfing on waves of sur- factant.Proc. Natl. Acad. Sci. U. S. A., 106:18109–18113,

  114. [131]

    Fauvart, P

    M. Fauvart, P. Phillips, D. Bachaspatimayum, N. Verstraeten, J. Fransaer, J. Michiels, and J. Vermant. Surface tension gradient control of bacterial swarming in colonies ofPseu- domonas aeruginosa.Soft Matter, 8:70–76, 2012. doi: 10.1039/c1sm06002c

  115. [132]

    Trinschek, K

    S. Trinschek, K. John, and U. Thiele. Modelling of surfactant- driven front instabilities in spreading bacterial colonies.Soft Matter, 14:4464–4476, 2018. doi: 10.1039/c8sm00422f

  116. [133]

    Matsuyama and M

    T. Matsuyama and M. Matsushita. Fractal morphogenesis by a bacterial-cell population.Crit. Rev. Microbiol., 19:117–135,

  117. [134]

    Seminara, T

    A. Seminara, T. E. Angelini, J. N. Wilking, H. Vlamakis, S. Ebrahim, R. Kolter, D. A. Weitz, and M. P. Brenner. Os- motic spreading ofBacillus subtilisbiofilms driven by an ex- tracellular matrix.Proc. Natl. Acad. Sci. U. S. A., 109:1116– 1121, 2012. doi: 10.1073/pnas.1109261108

  118. [135]

    Ben-Jacob, I

    E. Ben-Jacob, I. Cohen, and H. Levine. Cooperative self- organization of microorganisms.Adv. Phys., 49:395–554,

  119. [136]

    J. Yan, C. D. Nadell, H. A. Stone, N. S. Wingreen, and B. L. Bassler. Extracellular-matrix-mediated osmotic pres- sure drivesvibrio choleraebiofilm expansion and cheater exclusion.Nat. Commun., 8:327, 2017. doi: 10.1038/ s41467-017-00401-1

  120. [137]

    Trinschek, K

    S. Trinschek, K. John, S. Lecuyer, and U. Thiele. Continuous vs. arrested spreading of biofilms at solid-gas interfaces - the role of surface forces.Phys. Rev. Lett., 119:078003, 2017. doi: 10.1103/PhysRevLett.119.078003

  121. [138]

    A. Cont, T. Rossy, Z. Al-Mayyah, and A. Persat. Biofilms deform soft surfaces and disrupt epithelia.eLife, 9:e56533,

  122. [139]

    doi: 10.1073/pnas.0905890106

  123. [140]

    Pietz, K

    A. Pietz, K. John, and U. Thiele. The role of substrate mechan- ics in osmotic biofilm spreading.Soft Matter, 21:2935–2945,

  124. [141]

    Faiza, R

    N. Faiza, R. Welch, and A. Patteson. Substrate stiffness modu- lates collective colony expansion of the social bacteriumMyx- ococcus xanthus.APL Bioeng., 9, 2025. doi: 10.1063/5. 0226619

  125. [143]

    Beltrame, E

    P. Beltrame, E. Knobloch, P. H¨anggi, and U. Thiele. Rayleigh and depinning instabilities of forced liquid ridges on hetero- geneous substrates.Phys. Rev. E, 83:016305, 2011. doi: 10.1103/PhysRevE.83.016305

  126. [144]

    Srinivasan, C

    S. Srinivasan, C. N. Kaplan, and L. Mahadevan. A multiphase theory for spreading microbial swarms and films.eLife, 8: e42697, 2019. doi: 10.7554/eLife.42697

  127. [145]

    C. P. Brangwynne, C. R. Eckmann, D. S. Courson, A. Ry- barska, C. Hoege, J. Gharakhani, F. J ¨ulicher, and A. A. Hy- man. Germline P granules are liquid droplets that localize by controlled dissolution/condensation.Science, 324:1729–1732,

  128. [146]

    Mangiarotti, N

    A. Mangiarotti, N. N. Chen, Z. L. Zhao, R. Lipowsky, and R. Dimova. Wetting and complex remodeling of membranes by biomolecular condensates.Nat. Commun., 14:2809, 2023. doi: 10.1038/s41467-023-37955-2

  129. [147]

    Liese, X

    S. Liese, X. P. Zhao, C. A. Weber, and F. J ¨ulicher. Chem- ically active wetting.Proc. Natl. Acad. Sci. U. S. A., 122: 2403083122, 2024. doi: 10.1073/pnas.2403083122

  130. [148]

    Goychuk, L

    A. Goychuk, L. Demarchi, I. Maryshev, and E. Frey. Self- consistent sharp interface theory of active condensate dy- namics.Phys. Rev. Res., 6:033082, 2024. doi: 10.1103/ PhysRevResearch.6.033082

  131. [149]

    M. E. Asp, M.-T. Ho Thanh, D. A. Germann, R. J. Carroll, A. Franceski, R. D. Welch, A. Gopinath, and A. E. Patte- son. Spreading rates of bacterial colonies depend on sub- strate stiffness and permeability.PNAS Nexus, 1, 2022. doi: 10.1093/pnasnexus/pgac025

  132. [150]

    M. E. Cates and J. Tailleur. Motility-induced phase separation. Annu. Rev. Condens. Matter Phys., 6:219–244, 2015. doi: 10. 1146/annurev-conmatphys-031214-014710

  133. [151]

    doi: 10.1039/D4SM01463D

  134. [152]

    Turci, R

    F. Turci, R. L. Jack, and N. B. Wilding. Partial and complete wetting of droplets of active Brownian particles.Soft Matter, 20:2060–2074, 2024. doi: 10.1039/d3{s}m01493{b}

  135. [153]

    Hennes, J

    M. Hennes, J. Tailleur, G. Charron, and A. Daerr. Active de- pinning of bacterial droplets: the collective surfing ofbacillus subtilis.Proc. Natl. Acad. Sci. U. S. A., 114:5958–5963, 2017. doi: 10.1073/pnas.1703997114

  136. [154]

    Sep ´ulveda and R

    N. Sep ´ulveda and R. Soto. Universality of active wetting transitions.Phys. Rev. E, 98:052141, 2018. doi: 10.1103/ PhysRevE.98.052141

  137. [155]

    Berry, C

    J. Berry, C. P. Brangwynne, and M. Haataja. Physical princi- ples of intracellular organization via active and passive phase transitions.Rep. Prog. Phys., 80:046601, 2018. doi: 10.1088/ 1361-6633/aaa61e

  138. [156]

    Wittmann and J

    R. Wittmann and J. M. Brader. Active Brownian particles at in- terfaces: An effective equilibrium approach.Europhys. Lett., 114:68004, 2016. doi: 10.1209/0295-5075/114/68004

  139. [157]

    doi: 10.1126/science.1172046

  140. [158]

    Rojas-Vega, P

    M. Rojas-Vega, P. De Castro, and R. Soto. Mixtures of self-propelled particles interacting with asymmetric obsta- cles.Eur. Phys. J. E, 46:95, 2023. doi: 10.1140/epje/ s10189-023-00354-y

  141. [159]

    Rojas-Vega, P

    M. Rojas-Vega, P. de Castro, and R. Soto. Wetting dy- namics by mixtures of fast and slow self-propelled particles. Phys. Rev. E, 107:014608, 2023. doi: 10.1103/physreve.107. 014608

  142. [160]

    Caprini, D

    L. Caprini, D. Breoni, A. Ldov, C. Scholz, and H. L ¨owen. Dynamical clustering and wetting phenomena in inertial ac- tive matter.Commun. Phys., 7:343, 2024. doi: 10.1038/ s42005-024-01835-y

  143. [161]

    P. D. Neta, M. Tasinkevych, M. M. Telo da Gama, and C. S. Dias. Wetting of a solid surface by active matter.Soft Matter, 17:2468–2478, 2021. doi: 10.1039/d0{s}m02008g

  144. [162]

    Y . Zhao, R. Zakine, A. Daerr, Y . Kafri, J. Tailleur, and F. Van Wijland. Active Young-Dupr ´e equation: How self- organized currents stabilize partial wetting. 2024. doi: 10.48550/ARXIV .2405.20651

  145. [163]

    Turci and N

    F. Turci and N. B. Wilding. Wetting transition of active Brow- Preprint– contact: u.thiele@uni-muenster.de – www.uwethiele.de – February 12, 2026 12 nian particles on a thin membrane.Physical Review Letters, 127:238002, 2021. doi: 10.1103/physrevlett.127.238002

  146. [164]

    Wysocki and H

    A. Wysocki and H. Rieger. Capillary action in scalar active matter.Phys. Rev. Lett., 124:048001, 2020. doi: 10.1103/ PhysRevLett.124.048001

  147. [165]

    Sep ´ulveda and R

    N. Sep ´ulveda and R. Soto. Wetting transitions displayed by persistent active particles.Phys. Rev. Lett., 119:078001, 2017. doi: 10.1103/physrevlett.119.078001

  148. [166]

    Adkins, I

    R. Adkins, I. Kolvin, Z. H. You, S. Witthaus, M. C. Marchetti, and Z. Dogic. Dynamics of active liquid interfaces.Science, 377:768–772, 2022. doi: 10.1126/science.abo5423

  149. [167]

    Perez-Bast´ıas and R

    P. Perez-Bast´ıas and R. Soto. Two-field theory for phase co- existence of active Brownian particles. 2025. doi: 10.48550/ ARXIV .2504.13327

  150. [168]

    Kruse, J

    K. Kruse, J. F. Joanny, F. J ¨ulicher, J. Prost, and K. Seki- moto. Asters, vortices, and rotating spirals in active gels of polar filaments.Phys. Rev. Lett., 92:078101, 2004. doi: 10.1103/PhysRevLett.92.078101

  151. [169]

    Das and R

    S. Das and R. Chelakkot. Active wetting transitions induced by rotational noise at solid interfaces.J. Chem. Phys., 163: 014704, 2025. doi: 10.1063/5.0272268

  152. [170]

    Tjhung, D

    E. Tjhung, D. Marenduzzo, and M. E. Cates. Spontaneous symmetry breaking in active droplets provides a generic route to motility.Proc. Natl. Acad. Sci. U. S. A., 109(31):12381– 12386, 2012. doi: 10.1073/pnas.1200843109

  153. [171]

    Tjhung, A

    E. Tjhung, A. Tiribocchi, D. Marenduzzo, and M. E. Cates. A minimal physical model captures the shapes of crawling cells. Nat. Commun., 6, 2015. doi: 10.1038/ncomms6420

  154. [172]

    Khoromskaia and G

    D. Khoromskaia and G. P. Alexander. Motility of active fluid drops on surfaces.Phys. Rev. E, 92:062311, 2015. doi: 10. 1103/PhysRevE.92.062311

  155. [173]

    Grodzinski, R

    N. Grodzinski, R. L. Jack, and M. E. Cates. Hydrodynamic theory of wetting by active particles. June 2025. doi: 10. 48550/ARXIV .2506.14559

  156. [174]

    Loisy, J

    A. Loisy, J. Eggers, and T. B. Liverpool. Tractionless self- propulsion of active drops.Phys. Rev. Lett., 123:248006, 2019. doi: 10.1103/PhysRevLett.123.248006

  157. [175]

    Solon and Y

    A. Solon and Y . Zhao. The surprising physics of interfaces in active matter.Chinese Phys. Lett., 42:100901, 2025. doi: 10.1088/0256-307x/42/10/100901

  158. [176]

    Trinschek, F

    S. Trinschek, F. Stegemerten, K. John, and U. Thiele. Thin- film modelling of resting and moving active droplets.Phys. Rev. E, 101:062802, 2020. doi: 10.1103/PhysRevE.101. 062802

  159. [177]

    Mangeat, S

    M. Mangeat, S. Chakraborty, A. Wysocki, and H. Rieger. Stationary particle currents in sedimenting active matter wet- ting a wall.Phys. Rev. E, 109:014616, 2024. doi: 10.1103/ PhysRevE.109.014616

  160. [178]

    R. C. V . Coelho, H. R. J. C. Figueiredo, and M. M. Telo da Gama. Active nematics on flat surfaces: From droplet motility and scission to active wetting.Phys. Rev. Research, 5:033165,

  161. [179]

    J. F. Joanny and S. Ramaswamy. A drop of active matter.J. Fluid Mech., 705:46–57, 2012. doi: 10.1017/jfm.2012.131

  162. [180]

    G. R. Chandel and S. Das. Theory of soft active equilibrium wetting.J. Fluid Mech., 1019:A39, 2025. doi: 10.1017/jfm. 2025.10624

  163. [181]

    J. M. Oliver, J. R. King, K. J. McKinlay, P. D. Brown, D. M. Grant, C. A. Scotchford, and J. V . Wood. Thin-film theo- ries for two-phase reactive flow models of active cell motion. Math. Med. Biol., 22:53–98, 2005. doi: 10.1093/imammb/ dqh022

  164. [182]

    Z. H. You, A. Baskaran, and M. C. Marchetti. Nonreciproc- ity as a generic route to traveling states.Proc. Natl. Acad. Sci. U. S. A., 117:19767–19772, 2020. doi: 10.1073/pnas. 2010318117

  165. [183]

    Frohoff-H ¨ulsmann, J

    T. Frohoff-H ¨ulsmann, J. Wrembel, and U. Thiele. Suppres- sion of coarsening and emergence of oscillatory behavior in a Cahn-Hilliard model with nonvariational coupling.Phys. Rev. E, 103:042602, 2021. doi: 10.1103/PhysRevE.103.042602

  166. [184]

    Dinelli, J

    A. Dinelli, J. O’Byrne, A. Curatolo, Y . Zhao, P. Sollich, and J. Tailleur. Non-reciprocity across scales in active mixtures. Nat. Commun., 14, 2023. doi: 10.1038/s41467-023-42713-5

  167. [185]

    C. A. Whitfield and R. J. Hawkins. Instabilities, motion and deformation of active fluid droplets.New J. Phys., 18:123016,

  168. [186]

    doi: 10.1088/1367-2630/18/12/123016

  169. [187]

    Evans and A

    D. Evans and A. K. Omar. Theory of nonequilibrium coexis- tence with coupled conserved and nonconserved order param- eters.Phys. Rev. Res., 7, 2025. doi: 10.1103/2jgf-yb82

  170. [188]

    Loisy, J

    A. Loisy, J. Eggers, and T. B. Liverpool. How many ways a cell can move: the modes of self-propulsion of an active drop. Soft Matter, 16:3106–3124, 2020. doi: 10.1039/d0sm00070a

  171. [189]

    Greve, G

    D. Greve, G. Lovato, T. Frohoff-H ¨ulsmann, and U. Thiele. Coexistence of uniform and oscillatory states resulting from nonreciprocity and conservation laws.Phys. Rev. Lett., 134: 018303, 2025. doi: 10.1103/PhysRevLett.134.018303

  172. [190]

    Stegemerten, K

    F. Stegemerten, K. John, and U. Thiele. Symmetry-breaking, motion and bistability of active drops through polarization- surface coupling.Soft Matter, 18:5823–5832, 2022. doi: 10.1039/D2SM00648K

  173. [191]

    Weyer, T

    H. Weyer, T. A. Roth, and E. Frey. Deciphering the interface laws of Turing mixtures and foams. 2024. doi: 10.48550/ arxiv.2409.20070

  174. [192]

    A. K. Omar, K. Klymko, T. GrandPre, and P. L. Geissler. Phase diagram of active Brownian spheres: crystallization and the metastability of motility-induced phase separation.Phys. Rev. Lett., 126:188002, 2021. doi: 10.1103/PhysRevLett.126. 188002

  175. [193]

    Y . K. Li and T. Z. Qian. Hydrodynamics of a thin film of active nematic fluid: stationary state, spreading, and migration.Phys. Fluids, 37:072109, 2025. doi: 10.1063/5.0276281

  176. [194]

    John and U

    K. John and U. Thiele. Liquid transport generated by a flash- ing field-induced wettability ratchet.Appl. Phys. Lett., 90: 264102, 2007. doi: 10.1063/1.2751582

  177. [195]

    S. Saha, J. Agudo-Canalejo, and R. Golestanian. Scalar active mixtures: The non-reciprocal Cahn-Hilliard model.Phys. Rev. X, 10:041009, 2020. doi: 10.1103/PhysRevX.10.041009

  178. [196]

    Stieneker, L

    M. Stieneker, L. Topp, S. V . Gurevich, and A. Heuer. Multi- scale perspective on wetting on switchable substrates: map- ping between microscopic and mesoscopic models.Phys. Rev. Fluids, 8:013902, 2023. doi: 10.1103/PhysRevFluids. 8.013902

  179. [197]

    Mugele, A

    F. Mugele, A. Klingner, J. Buehrle, D. Steinhauser, and S. Her- minghaus. Electrowetting: a convenient way to switchable wettability patterns.J. Phys.: Condens. Matter, 17:S559– S576, 2005. doi: 10.1088/0953-8984/17/9/016

  180. [198]

    Zamboni, D

    R. Zamboni, D. Ray, C. Denz, and J. Imbrock. Optoelectric- driven wetting transition on artificially micropatterned sur- faces with long-range virtual electrodes.Adv. Mater. Inter- faces, 2024. doi: 10.1002/admi.202400459

  181. [199]

    Y .-J. Chiu, D. Evans, and A. K. Omar. Theory of nonequilib- rium multicomponent coexistence. 2024. doi: 10.48550/arxiv. 2409.07620

  182. [200]

    Brauns and M

    F. Brauns and M. C. Marchetti. Nonreciprocal pattern forma- tion of conserved fields.Phys. Rev. X, 14(2):021014, 2024. doi: 10.1103/physrevx.14.021014

  183. [201]

    Golomb, N

    M. Golomb, N. B. Arndt, C. Honnigfort, B. Shakhayeva, B. J. Ravoo, and B. Braunschweig. Molecular kinetics and wet- ting dynamics of self-assembled monolayers with fluorinated arylazopyrazoles.J. Phys. Chem. C, 127:15316–15325, 2023. doi: 10.1021/acs.jpcc.3c02472

  184. [202]

    Y . Duan, J. Agudo-Canalejo, R. Golestanian, and B. Ma- hault. Phase coexistence in nonreciprocal quorum-sensing ac- tive matter.Phys. Rev. Res., 7:013234, 2025. doi: 10.1103/ PhysRevResearch.7.013234. Preprint– contact: u.thiele@uni-muenster.de – www.uwethiele.de – February 1...

  185. [203]

    Dinelli, J

    A. Dinelli, J. O’Byrne, and J. Tailleur. Fluctuating hydrody- namics of active particles interacting via chemotaxis and quo- rum sensing: static and dynamics.J. Phys. A: Math. Theor., 57:395002, 2024. doi: 10.1088/1751-8121/ad72bc. Preprint– contact: u.thiele@uni-muenster.de –...

  186. [204]

    Ma and M

    X. Ma and M. E. Cates. Wetting and pattern formation in non- reciprocal ternary phase separation.New J. Phys., 27:124401,

  187. [205]

    doi: 10.1088/1367-2630/ae2883

  188. [208]

    M. P. Holl, A. B. Steinberg, M. te Vrugt, and U. Thiele. Motility-induced crystallization and rotating crystallites. Phys. Rev. Lett., 135:158301, 2025. doi: 10.1103/m3dy-53yc

  189. [210]

    Steering droplets on sub- strates using moving steps in wettability.Soft Matter, 17: 2454–2467, 2021

    Josua Grawitter and Holger Stark. Steering droplets on sub- strates using moving steps in wettability.Soft Matter, 17: 2454–2467, 2021. doi: 10.1039/d0sm02082f

  190. [214]

    Honnigfort, L

    C. Honnigfort, L. Topp, N. G. Rey, A. Heuer, and B. Braun- schweig. Dynamic wetting of photoresponsive arylazopyra- zole monolayers is controlled by the molecular kinetics of the monolayer.J. Am. Chem. Soc., 144:4026–4038, 2022. doi: 10.1021/jacs.1c12832

  191. [215]

    Nekoonam, G

    N. Nekoonam, G. Vera, A. Goralczyk, F. Mayoussi, P. Zhu, D. B ¨ocherer, A. Shakeel, and D. Helmer. Controllable wet- ting transitions on photoswitchable physical gels.ACS Appl. Mater. Interfaces, 15:27234–27242, 2023. ISSN 1944-8252. doi: 10.1021/acsami.2c22979

  192. [217]

    Avanzini, T

    F. Avanzini, T. Aslyamov, ´E. Fodor, and M. Espos- ito. Nonequilibrium thermodynamics of non-ideal re- action–diffusion systems: Implications for active self- organization.J. Chem. Phys., 161:174108, 2024. doi: 10. 1063/5.0231520

  193. [1993]

    doi: 10.3109/10408419309113526

  194. [2000]

    doi: 10.1080/000187300405228

  195. [2009]

    doi: 10.1140/epjst/e2009-00902-3

  196. [2016]

    doi: 10.1007/s10853-015-9331-3

  197. [2017]

    doi: 10.1016/j.eml.2017.01.004

  198. [2020]

    doi: 10.7554/elife.56533

  199. [2023]

    doi: 10.1103/physrevresearch.5

    ISSN 2643-1564. doi: 10.1103/physrevresearch.5. 033165

  200. [2025]

    doi: 10.1103/f3ck-dx5c

Pith tools

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