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Paper Citation Record · LEDGER

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond

As of 9 August 2026, this Paper Citation Record lists 54 of 54 outbound references and 0 inbound Pith citation observations for arXiv:2510.02853.

A citation records a reference. It does not transfer a finding from one paper to another.

pith.paper-citation-record.v1
2510.02853 v3

Coverage vector

measured 54 of 54 reference resolution

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Source: paper_references, paper_reference_links, observed 2026-08-04T12:44:52.165988Z

measured 54 of 54 standing notices

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54 of 54 outbound references displayed

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Outbound references

Observation 8275d2f1-7d92-48d1-b06e-bfd196520304 · outbound

This paper cites From the neuron doctrine to neural networks.Nature reviews neuroscience, 16(8):487–497, 2015.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond From the neuron doctrine to neural networks.Nature reviews neuroscience, 16(8):487–497, 2015

Reference 1

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Observation 2c554a4b-8268-4e1e-b953-0566ad3010ae · outbound

This paper cites Computation through neural population dynamics.Annual review of neuroscience, 43(1):249–275, 2020.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Computation through neural population dynamics.Annual review of neuroscience, 43(1):249–275, 2020

Reference 2

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Observation ab978ea6-b2d3-4c27-8998-8baa82b5210a · outbound

This paper cites Neural tuning and representational geometry.Nature Reviews Neuroscience, 22(11):703–718, 2021.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Neural tuning and representational geometry.Nature Reviews Neuroscience, 22(11):703–718, 2021

Reference 3

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Observation 9962b709-2ca4-4260-ace3-23973f3f4095 · outbound

This paper cites Aneuralmanifoldviewofthebrain.NatureNeuroscience, pages 1–16, 2025.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Aneuralmanifoldviewofthebrain.NatureNeuroscience, pages 1–16, 2025

Reference 4

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Observation 1be1b993-3e74-480b-9a6f-047733b40d90 · outbound

This paper cites Large-scale neural recordings call for new insights to link brain and behavior.Nature neuroscience, 25(1):11–19, 2022.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Large-scale neural recordings call for new insights to link brain and behavior.Nature neuroscience, 25(1):11–19, 2022

Reference 5

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Observation 18467938-d9fa-487d-9216-08fd120a872d · outbound

This paper cites Neural population dynamics during reaching.Nature, 487(7405):51–56, 2012.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Neural population dynamics during reaching.Nature, 487(7405):51–56, 2012

Reference 6

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Observation 3dfce388-30b2-448c-aebd-0f605f25a2f0 · outbound

This paper cites Towardstheneuralpopulationdoctrine.Currentopinioninneurobiology, 55:103–111, 2019.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Towardstheneuralpopulationdoctrine.Currentopinioninneurobiology, 55:103–111, 2019

Reference 7

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Observation 8c38a751-f79c-45be-afb3-b8d49ff0f251 · outbound

This paper cites A unifying perspective on neural manifolds and circuits for cognition.Nature Reviews Neuroscience, 24(6):363–377, 2023.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond A unifying perspective on neural manifolds and circuits for cognition.Nature Reviews Neuroscience, 24(6):363–377, 2023

Reference 8

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Observation 0db61b56-b2d3-46da-93d5-dce15e9cbaea · outbound

This paper cites Spatial representation in the entorhinal cortex.Science, 305(5688):1258–1264, 2004.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Spatial representation in the entorhinal cortex.Science, 305(5688):1258–1264, 2004

Reference 9

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Observation b8e4860c-e4bd-4130-976b-b1d239045200 · outbound

This paper cites Microstructure of a spatial map in the entorhinal cortex.Nature, 436(7052):801–806, 2005.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Microstructure of a spatial map in the entorhinal cortex.Nature, 436(7052):801–806, 2005

Reference 10

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Observation fb4253ee-9891-493c-b405-b2aae8db0d2d · outbound

This paper cites Conjunctive representation of position, direction, and velocity in entorhinal cortex.Science, 312(5774):758–762, 2006.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Conjunctive representation of position, direction, and velocity in entorhinal cortex.Science, 312(5774):758–762, 2006

Reference 11

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Observation 4c9cde59-6a1b-4812-985d-cfea2b016afc · outbound

This paper cites Toroidal topology of population activity in grid cells.Nature, 602(7895):123–128, 2022.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Toroidal topology of population activity in grid cells.Nature, 602(7895):123–128, 2022

Reference 12

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Observation 1a655883-bc8c-4a15-9a35-33204ed8d253 · outbound

This paper cites Path integration and the neural basis of the’cognitive map’.Nature Reviews Neuroscience, 7(8):663–678, 2006.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Path integration and the neural basis of the’cognitive map’.Nature Reviews Neuroscience, 7(8):663–678, 2006

Reference 13

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Observation 9722a2f2-28a3-4f89-ad23-f0f55b666505 · outbound

This paper cites Accuratepathintegrationincontinuousattractornetworkmodelsofgridcells.PLoS computational biology, 5(2):e1000291, 2009.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Accuratepathintegrationincontinuousattractornetworkmodelsofgridcells.PLoS computational biology, 5(2):e1000291, 2009

Reference 14

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Observation 6638146f-53ec-44b2-9b8a-a57d78148c7a · outbound

This paper cites Impairedpathintegrationinmicewithdisruptedgridcellfiring.Natureneuroscience,21(1):81–91,2018.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Impairedpathintegrationinmicewithdisruptedgridcellfiring.Natureneuroscience,21(1):81–91,2018

Reference 15

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Observation bd4c5522-414b-446c-be1c-5b6e1b381202 · outbound

This paper cites A metric for space.Hippocampus, 18(12):1142–1156, 2008.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond A metric for space.Hippocampus, 18(12):1142–1156, 2008

Reference 16

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Observation 27e25033-3fd7-4c46-934e-5501d7ce020a · outbound

This paper cites Are grid cells used for navigation? on local metrics, subjective spaces, and black holes.Neuron, 111(12):1858–1875, 2023.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Are grid cells used for navigation? on local metrics, subjective spaces, and black holes.Neuron, 111(12):1858–1875, 2023

Reference 17

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Observation f6112475-ef4b-45c7-8e48-3033f67e563a · outbound

This paper cites The hippocampus as a predictive map.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond The hippocampus as a predictive map

Reference 18

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Observation e7bb5a99-8201-44ad-a6fb-df7da4c5a24a · outbound

This paper cites Vector-based navigation using grid-like representations in artificial agents.Nature, 557(7705):429–433, 2018.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Vector-based navigation using grid-like representations in artificial agents.Nature, 557(7705):429–433, 2018

Reference 19

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Observation 1ca22d36-e865-4611-8d06-5d81d616614a · outbound

This paper cites Emergence of grid-like representations by training recurrent neural networks to perform spatial localization.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Emergence of grid-like representations by training recurrent neural networks to perform spatial localization

Reference 20

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Observation eb5005cc-4d11-4edc-9f9f-18b8e1b68f9a · outbound

This paper cites A unified theory for the origin of grid cells through the lens of pattern formation.Advances in neural information processing systems, 32, 2019.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond A unified theory for the origin of grid cells through the lens of pattern formation.Advances in neural information processing systems, 32, 2019

Reference 21

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Observation 1a15939e-c4a2-49e9-a0ea-ad2caea692c3 · outbound

This paper cites The tolman-eichenbaum machine: unifying space and relational memory through generalization in the hippocampal formation.Cell, 183(5):1249–1263, 2020.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond The tolman-eichenbaum machine: unifying space and relational memory through generalization in the hippocampal formation.Cell, 183(5):1249–1263, 2020

Reference 22

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Observation 9866accc-e327-45bd-b7e6-47a3ec812706 · outbound

This paper cites On path integration of grid cells: group representation and isotropic scaling.Advances in Neural Information Processing Systems, 34:28623–28635, 2021.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond On path integration of grid cells: group representation and isotropic scaling.Advances in Neural Information Processing Systems, 34:28623–28635, 2021

Reference 23

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Observation 8ea7cedf-81a6-4ff9-afad-eec207ed6803 · outbound

This paper cites Conformal Isometry of Lie Group Representation in Recurrent Network of Grid Cells.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Conformal Isometry of Lie Group Representation in Recurrent Network of Grid Cells

Reference 24

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Observation 0623f463-79e3-4816-9dc4-942de130c5fb · outbound

This paper cites Actionable Neural Representations: Grid Cells from Minimal Constraints.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Actionable Neural Representations: Grid Cells from Minimal Constraints

Reference 25

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Observation 7bbdeb7c-911b-4563-9f4d-e15fcb49a7f0 · outbound

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The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Unresolved cited work

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Observation 04e4f306-a002-4a36-ad72-6b3958814799 · outbound

This paper cites Coherently remapping toroidal cells but not grid cells are responsible for path integration in virtual agents.Iscience, 26(11), 2023.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Coherently remapping toroidal cells but not grid cells are responsible for path integration in virtual agents.Iscience, 26(11), 2023

Reference 27

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Observation ae34ff93-8000-4a64-b05f-12e72a38faca · outbound

This paper cites A unified theory for the computational and mechanistic origins of grid cells.Neuron, 111(1):121–137, 2023.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond A unified theory for the computational and mechanistic origins of grid cells.Neuron, 111(1):121–137, 2023

Reference 28

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Observation ea00b5ac-9e68-4ffe-85c4-bdaa09b8d86b · outbound

This paper cites Self-supervised learningofrepresentationsforspacegeneratesmulti-modulargridcells.AdvancesinNeuralInformationProcessing Systems, 36:23140–23157, 2023.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Self-supervised learningofrepresentationsforspacegeneratesmulti-modulargridcells.AdvancesinNeuralInformationProcessing Systems, 36:23140–23157, 2023

Reference 29

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Observation c5317943-3892-4ef3-a0c2-d0d23373be26 · outbound

This paper cites Self-supervised grid cells without path integration.bioRxiv, pages 2024–05, 2024.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Self-supervised grid cells without path integration.bioRxiv, pages 2024–05, 2024

Reference 30

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Observation ef6a82a1-7e30-4f06-b21f-b97a4f9deb03 · outbound

This paper cites On conformal isometry of grid cells: Learning distance-preserving position embedding.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond On conformal isometry of grid cells: Learning distance-preserving position embedding

Reference 31

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Observation 1c734332-7f21-4baa-824f-0a46d27fcae3 · outbound

This paper cites Some informational aspects of visual perception.Psychological review, 61(3):183, 1954.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Some informational aspects of visual perception.Psychological review, 61(3):183, 1954

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Observation 230b0903-5bad-4744-aa44-7036181b252e · outbound

This paper cites Possible principles underlying the transformation of sensory messages.Sensory communication, 1(01):217–233, 1961.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Possible principles underlying the transformation of sensory messages.Sensory communication, 1(01):217–233, 1961

Reference 33

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Observation 56efdc15-e39b-4d78-9b51-b2eff7908b67 · outbound

This paper cites Second-order isomorphism of internal representations: Shapes of states.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Second-order isomorphism of internal representations: Shapes of states

Reference 34

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Observation 254d2b48-6cf0-419e-bcc8-ee83b4f6facc · outbound

This paper cites Representation is representation of similarities.Behavioral and brain sciences, 21(4):449–467, 1998.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Representation is representation of similarities.Behavioral and brain sciences, 21(4):449–467, 1998

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source=pdf_text observed=2026-08-04T12:44:50.860739Z digest=sha256:561e3b7510cb5144b06e18cdf8e2aab31acd6e527ea922d1e18e64a32d520036

Observation 1f566a23-614b-46af-a4c8-a49863195c7c · outbound

This paper cites No free lunch theorems for optimization.IEEE transactions on evolutionary computation, 1(1):67–82, 1997.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond No free lunch theorems for optimization.IEEE transactions on evolutionary computation, 1(1):67–82, 1997

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source=pdf_text observed=2026-08-04T12:44:50.925182Z digest=sha256:851450e0e5dafec44a1298f2425ec0ea7ec401a0f45af1efd78de43d5f571437

Observation db3e8fb6-8c35-4c4d-8d2f-a84b738a935e · outbound

This paper cites Representation learning: A review and new perspectives.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Representation learning: A review and new perspectives

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source=pdf_text observed=2026-08-04T12:44:51.052528Z digest=sha256:277d05452aa33d604a596046c37d4f0df70f91b564d0f125d5733559a2bac625

Observation 8378a77c-029c-42ea-819d-3e776ed113b8 · outbound

This paper cites Geometric Deep Learning: Grids, Groups, Graphs, Geodesics, and Gauges.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Geometric Deep Learning: Grids, Groups, Graphs, Geodesics, and Gauges

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source=pdf_text observed=2026-08-04T12:44:51.105889Z digest=sha256:5590c5df43e5155761792ae7b1d623663c9a86888093f5b81973e116ab4c0d41

Observation 78de2de9-28ed-450b-bba6-09dd69393a2f · outbound

This paper cites Getting aligned on representational alignment.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Getting aligned on representational alignment

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source=pdf_text observed=2026-08-04T12:44:51.196076Z digest=sha256:879581ff2739952696de03a22fee6beda0420a5537665a4fe0e87c40bc7fd2e6

Observation fe6e8577-ec1d-4f16-88b0-d1275204816e · outbound

This paper cites What grid cells convey about rat location.Journal of Neuroscience, 28(27):6858–6871, 2008.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond What grid cells convey about rat location.Journal of Neuroscience, 28(27):6858–6871, 2008

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source=pdf_text observed=2026-08-04T12:44:51.267970Z digest=sha256:113428b9549f6b32355019834032f6d550862e178099d6a35809e28bd36f833d

Observation 3926f411-1180-4f04-bd63-3f1dd0aadc2e · outbound

This paper cites PrincetonUniversity Press, 2021.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond PrincetonUniversity Press, 2021

Reference 41

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source=pdf_text observed=2026-08-04T12:44:51.335783Z digest=sha256:b714154412f45539ed1dadcc8d447ab64f1fee30ce043cdf728ff6768cff4e4a

Observation a2e0aa64-b91c-4d88-9258-0bddc003ae40 · outbound

This paper cites Cambridge University Press, 2002.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Cambridge University Press, 2002

Reference 42

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source=pdf_text observed=2026-08-04T12:44:51.401538Z digest=sha256:63d26fda1d0e59a4c58a8f73385a2606a548aa05c0c61e090d0c9bc9ec8ee0f3

Observation 34563d37-aaf0-4a2e-b83a-73589bcdc9ac · outbound

This paper cites Grid cells generate an analog error-correcting code for singularly precise neural computation.Nature neuroscience, 14(10):1330–1337, 2011.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Grid cells generate an analog error-correcting code for singularly precise neural computation.Nature neuroscience, 14(10):1330–1337, 2011

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source=pdf_text observed=2026-08-04T12:44:51.445382Z digest=sha256:80318e63c34cff2f9b9ff73b8f0f43e2e22a60af7e858e248166f97f5db8395d

Observation 6b0efabc-70d9-464d-8cc1-ccb15ae8ed29 · outbound

This paper cites The elementary geometric structure of compact lie groups.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond The elementary geometric structure of compact lie groups

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source=pdf_text observed=2026-08-04T12:44:51.467518Z digest=sha256:9ccd76ba0b6de599b90e245ab199c47f1064ed2f61bc91274f3f7e1f91a34e28

Observation 836e3beb-80dd-45e5-a7c2-51131997489a · outbound

This paper cites Head-directioncellsinthedeeplayersofdorsalpresubiculumoffreelymovingrats.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Head-directioncellsinthedeeplayersofdorsalpresubiculumoffreelymovingrats

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source=pdf_text observed=2026-08-04T12:44:51.533710Z digest=sha256:434e59c10aa6922a70043d78cb836459704262b0e269cb758480517cff27876f

Observation 17c5aa5d-7ede-42e0-80be-77d739dc9a37 · outbound

This paper cites Locally ordered representation of 3d space in the entorhinal cortex.Nature, 596(7872):404–409, 2021.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Locally ordered representation of 3d space in the entorhinal cortex.Nature, 596(7872):404–409, 2021

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source=pdf_text observed=2026-08-04T12:44:51.622164Z digest=sha256:5c8ef2f6e2d5f41b8f5377e11b2d3642e2ee9e7df60df6506fffde0b15054e51

Observation 0d89e736-2b5d-4515-8ffd-938f1464eee1 · outbound

This paper cites Irregular distribution of grid cell firing fields in rats exploring a 3d volumetric space.Nature neuroscience, 24(11):1567–1573, 2021.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Irregular distribution of grid cell firing fields in rats exploring a 3d volumetric space.Nature neuroscience, 24(11):1567–1573, 2021

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source=pdf_text observed=2026-08-04T12:44:51.682020Z digest=sha256:77839c48f13fdea4624d2c892725ee3194416042edbab6b0a9c0a0d8898f658b

Observation 8a2a9158-7610-4f27-86c9-4b3e153750e9 · outbound

This paper cites A model of the neural basis of the rat’s sense of direction.Advances in neural information processing systems, 7, 1994.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond A model of the neural basis of the rat’s sense of direction.Advances in neural information processing systems, 7, 1994

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source=pdf_text observed=2026-08-04T12:44:51.800531Z digest=sha256:ad03cbb881fc8313ebfea79a13f6bfe9bd1e311b456d0db8bdb2554beced026e

Observation 89208220-91cf-4056-8779-67d058adf228 · outbound

This paper cites Representation of spatial orientation by the intrinsic dynamics of the head-direction cell ensemble: a theory.Journal of Neuroscience, 16(6):2112–2126, 1996.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Representation of spatial orientation by the intrinsic dynamics of the head-direction cell ensemble: a theory.Journal of Neuroscience, 16(6):2112–2126, 1996

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source=pdf_text observed=2026-08-04T12:44:51.889252Z digest=sha256:238c57eb3a625a991754f6d3550b90aa37db95bdafc3c2e9468b32baabab6e09

Observation 81eecba8-bbc2-4e18-83e6-690e41503262 · outbound

This paper cites Grid cell symmetry is shaped by environmental geometry.Nature, 518(7538):232–235, 2015.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Grid cell symmetry is shaped by environmental geometry.Nature, 518(7538):232–235, 2015

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source=pdf_text observed=2026-08-04T12:44:51.943457Z digest=sha256:ef70ca05ea2f39453cc65f7d57a375a906ae30826f6831ad095291ae37b0cbb8

Observation 7d9699ea-172e-43a0-a8eb-61b3f72fb3b6 · outbound

This paper cites Local transformations of the hippocampal cognitive map.Science, 359(6380):1143–1146, 2018.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Local transformations of the hippocampal cognitive map.Science, 359(6380):1143–1146, 2018

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source=pdf_text observed=2026-08-04T12:44:52.027294Z digest=sha256:140fcfc6f7d9151975328351b99e5f3590353a0848f6fcbb9f2714616f966214

Observation 7bedaf61-a424-4309-b82e-05cd2a4f43b4 · outbound

This paper cites Shearing-induced asymmetry in entorhinal grid cells.Nature, 518(7538):207–212, 2015.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Shearing-induced asymmetry in entorhinal grid cells.Nature, 518(7538):207–212, 2015

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source=pdf_text observed=2026-08-04T12:44:52.077549Z digest=sha256:bf551f92ca36e15ec269094db3ecd0c11ce234ebda9ec992f67266da4d886656

Observation 64b955ac-8ae7-4919-80d7-c233b4483c06 · outbound

This paper cites Information processing with population codes.Nature Reviews Neuroscience, 1(2):125–132, 2000.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Information processing with population codes.Nature Reviews Neuroscience, 1(2):125–132, 2000

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source=pdf_text observed=2026-08-04T12:44:52.135359Z digest=sha256:a74e4fde8eae0abce944fc524cef69eefb52386330fb8f802645b540e85ae5f2

Observation 270eacf7-4bdb-45a5-9f3d-8130495e96fd · outbound

This paper cites Connecting multiple spatial scales to decode the population activity of grid cells.Science Advances, 1(11):e1500816, 2015.

The Principle of Isomorphism: A Theory of Population Activity in Grid Cells and Beyond Connecting multiple spatial scales to decode the population activity of grid cells.Science Advances, 1(11):e1500816, 2015

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source=pdf_text observed=2026-08-04T12:44:52.165988Z digest=sha256:175ff9dd48d2c02e43208e03f00014eec6f7c5e05bce67d7131af8eda2b800f9

Pith citing papers

No inbound Pith citation observations are available.