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Paper Citation Record · LEDGER

DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

As of 9 August 2026, this Paper Citation Record lists 0 of 0 outbound references and 29 inbound Pith citation observations for arXiv:2306.15006.

A citation records a reference. It does not transfer a finding from one paper to another.

pith.paper-citation-record.v1
2306.15006 v2

Coverage vector

measured 0 of 0 reference resolution

Typed states for the displayed outbound observations.

Source: paper_references, paper_reference_links

measured 29 of 29 standing notices

One-hop event checks from named stored sources.

Source: scholarly_work_events, retraction_status_cache, observed 2026-08-09T06:31:02.800959+00:00

measured 29 of 29 inbound itemization

Pith citing papers itemized under the disclosed page cap.

Source: paper_references, paper_reference_links, observed 2026-08-09T10:16:49.313150Z

measured 0 of 1 external citation measurements

A source-named dated measurement, never combined with another source.

Source: arxiv_reference, observed 2026-07-03T03:57:38.576237Z

Reference resolution

0 of 0 outbound references displayed

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External citation measurements

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Outbound references

No outbound reference observations are available for this paper version.

Pith citing papers

Observation d75c0912-fecb-4add-ad52-d63807033712 · inbound

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning cites this paper.

Omni-DNA: A Unified Genomic Foundation Model for Cross-Modal and Multi-Task Learning DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 53

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no resolver link, observed 2026-08-09T10:16:49.313150Z

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Unavailable: canonical work link unavailable.

source=arxiv_source observed=2026-08-09T10:16:49.313150Z digest=sha256:5558d2f466c44e7a3e06c568bbad73e2892be7d685858f87cd8f16fc9f2db359

Observation 3c572736-a75f-4926-922d-9de34d8471ce · inbound

Find Central Dogma Again: Leveraging Multilingual Transfer in Large Language Models cites this paper.

Find Central Dogma Again: Leveraging Multilingual Transfer in Large Language Models DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 12

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no resolver link, observed 2026-08-08T16:17:55.128254Z

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source=pdf_text observed=2026-08-08T16:17:55.128254Z digest=sha256:c2f01302e9f438e43dd5960bb9ce0a56eeeaf14690ae58a3f78688b312a80ae4

Observation 8db58757-dc8f-4cff-8697-db40984be141 · inbound

Nature Language Model: Deciphering the Language of Nature for Scientific Discovery cites this paper.

Nature Language Model: Deciphering the Language of Nature for Scientific Discovery DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 96

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no resolver link, observed 2026-08-08T12:31:52.298340Z

Source-reported events for the cited work

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source=pdf_text observed=2026-08-08T12:31:52.298340Z digest=sha256:51387c811a11edec2d5c67f6137b5acc628f0a830d4255ea99ca6863d11fa4b1

Observation 6da71cb3-f4aa-4b62-901a-ac15323f4a1d · inbound

OmniGenBench: A Modular Platform for Reproducible Genomic Foundation Models Benchmarking cites this paper.

OmniGenBench: A Modular Platform for Reproducible Genomic Foundation Models Benchmarking DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 11

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no resolver link, observed 2026-08-07T15:39:02.491956Z

Source-reported events for the cited work

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source=pdf_text observed=2026-08-07T15:39:02.491956Z digest=sha256:89d05dd8f26cd3385114708705e1f1bdd85db78a4a1cf961b1921abcb0d2d84c

Observation 700e4e8a-caf4-45ef-bfb2-347f4c249a7d · inbound

Minimalist Softmax Attention Provably Learns Constrained Boolean Functions cites this paper.

Minimalist Softmax Attention Provably Learns Constrained Boolean Functions DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 26

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no resolver link, observed 2026-08-07T14:21:01.955998Z

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source=pdf_text observed=2026-08-07T14:21:01.955998Z digest=sha256:bb4e52182b830699a28a7e0a9b231fd953c42803cc102146e6499050d8075e1b

Observation 23d35700-a73e-4104-8252-f38eb3b3bf17 · inbound

HAD: Hybrid Architecture Distillation Outperforms Teacher in Genomic Sequence Modeling cites this paper.

HAD: Hybrid Architecture Distillation Outperforms Teacher in Genomic Sequence Modeling DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 4

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no resolver link, observed 2026-08-07T13:53:40.729297Z

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Unavailable: canonical work link unavailable.

source=pdf_text observed=2026-08-07T13:53:40.729297Z digest=sha256:9739b8de7bc992c10440148b786dcbfba393f8e34920af1aee20344881beaac4

Observation 8862244a-006e-4a9d-a184-d59a348ab690 · inbound

GeneBreaker: Jailbreak Attacks against DNA Language Models with Pathogenicity Guidance cites this paper.

GeneBreaker: Jailbreak Attacks against DNA Language Models with Pathogenicity Guidance DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 81

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no resolver link, observed 2026-08-07T13:15:24.749565Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

source=pdf_text observed=2026-08-07T13:15:24.749565Z digest=sha256:ec3a17476c4db528314b45a0f6fdb9b8688980027e3dbf45fe111d12aefd1051

Observation d72606d8-f719-4b34-8f87-030383e8a8b6 · inbound

Leveraging Natural Language Processing to Unravel the Mystery of Life: A Review of NLP Approaches in Genomics, Transcriptomics, and Proteomics cites this paper.

Leveraging Natural Language Processing to Unravel the Mystery of Life: A Review of NLP Approaches in Genomics, Transcriptomics, and Proteomics DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 26

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no resolver link, observed 2026-08-07T11:31:19.045431Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

source=pdf_text observed=2026-08-07T11:31:19.045431Z digest=sha256:4636e41cbd61a8b26f35b5edca37dcc44f1a1378dac64ccf09249ad22b15416e

Observation 7b0a3283-ed0b-4b25-a95b-460161addb09 · inbound

Evaluating DNA function understanding in genomic language models using evolutionarily implausible sequences cites this paper.

Evaluating DNA function understanding in genomic language models using evolutionarily implausible sequences DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 32

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no resolver link, observed 2026-08-07T04:36:21.667026Z

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Unavailable: canonical work link unavailable.

source=pdf_text observed=2026-08-07T04:36:21.667026Z digest=sha256:9173ae531f5a68cfead238240c48ce16464e6e51c358083f63685d94f74e0cd0

Observation 0a639e47-ef46-44d1-955f-1ae710012c0a · inbound

BMFM-DNA: A SNP-aware DNA foundation model to capture variant effects cites this paper.

BMFM-DNA: A SNP-aware DNA foundation model to capture variant effects DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 13

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no resolver link, observed 2026-08-06T22:36:02.166688Z

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source=pdf_text observed=2026-08-06T22:36:02.166688Z digest=sha256:a7d752a03a196a9dab4f86e165c75f6a28f5c36b7ae7f796f974bb4f2506e4f9

Observation bb3b5c4d-0897-4e4f-a23c-b0455a1cef72 · inbound

Evaluation of Coding Schemes for Transformer-based Gene Sequence Modeling cites this paper.

Evaluation of Coding Schemes for Transformer-based Gene Sequence Modeling DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 2015

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no resolver link, observed 2026-08-06T15:47:06.344749Z

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source=pdf_text observed=2026-08-06T15:47:06.344749Z digest=sha256:ac58d0de9e3f8bc4dacbe21e884212796ab5b33a9abba2a371cbe00c41f2554a

Observation 1b3ad77e-0132-4532-86e3-98db3743104e · inbound

Fast and Scalable Gene Embedding Search: A Comparative Study of FAISS and ScaNN cites this paper.

Fast and Scalable Gene Embedding Search: A Comparative Study of FAISS and ScaNN DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 23

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no resolver link, observed 2026-08-06T15:05:25.176787Z

Source-reported events for the cited work

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source=arxiv_source observed=2026-08-06T15:05:25.176787Z digest=sha256:253748fd97c666084226e6507a5f1f0a2633ecf2ec5a69a12a4309461cae8cfa

Observation edd983a4-cc16-4e81-849e-bd03cee932df · inbound

EnTao-GPM: DNA Foundation Model for Predicting the Germline Pathogenic Mutations cites this paper.

EnTao-GPM: DNA Foundation Model for Predicting the Germline Pathogenic Mutations DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 6

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Unavailable: canonical work link unavailable.

source=pdf_text observed=2026-08-06T12:34:03.697041Z digest=sha256:a332b171cacaf7340174ada7247127aed05e33d06923defac166e459afdb36d6

Observation 1b0fc6c4-36c4-4b13-9102-e6a29e814b30 · inbound

NucEL: Single-Nucleotide ELECTRA-Style Genomic Pre-training for Efficient and Interpretable Representations cites this paper.

NucEL: Single-Nucleotide ELECTRA-Style Genomic Pre-training for Efficient and Interpretable Representations DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 22

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no resolver link, observed 2026-08-05T20:00:19.244272Z

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source=arxiv_source observed=2026-08-05T20:00:19.244272Z digest=sha256:9ca03edb34db3b62b9a98095b5a59ec23af9dc061e56875f1fe0418025ccf38e

Observation 388a6010-4f5a-4a0d-a732-03dd31d965a9 · inbound

Generative Artificial Intelligence in Bioinformatics: A Systematic Review of Models, Applications, and Methodological Advances cites this paper.

Generative Artificial Intelligence in Bioinformatics: A Systematic Review of Models, Applications, and Methodological Advances DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 19

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no resolver link, observed 2026-08-03T23:59:54.514934Z

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source=pdf_text observed=2026-08-03T23:59:54.514934Z digest=sha256:73da959343726b76204d5fa061bcb5e27f19aac3fa23759b4872c266c87e0036

Observation e49a4bf5-7ed9-41ef-ac17-9f32135b0ca4 · inbound

BioArc: Discovering Optimal Neural Architectures for Biological Foundation Models cites this paper.

BioArc: Discovering Optimal Neural Architectures for Biological Foundation Models DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 4054

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Unavailable: canonical work link unavailable.

source=pdf_text observed=2026-08-03T19:32:01.164181Z digest=sha256:d4cd4ed4208f4cbe7e1ca81b1454e4d4ec4d0a626246329c9b0ce64b2fca7d1f

Observation da62046b-d94c-4010-ae79-4ccd9afd3665 · inbound

Rethinking Genomic Modeling Through Optical Character Recognition cites this paper.

Rethinking Genomic Modeling Through Optical Character Recognition DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 51

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no resolver link, observed 2026-08-03T05:34:18.210942Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

source=arxiv_source observed=2026-08-03T05:34:18.210942Z digest=sha256:9cb243b3599831e3e5396d5ec5bc1853157726c7c4b8aeeb9da3421fa69d78c3

Observation 1246c614-d824-4f69-8608-a946486481ed · inbound

JEPA-DNA: Grounding Genomic Foundation Models through Joint-Embedding Predictive Architectures cites this paper.

JEPA-DNA: Grounding Genomic Foundation Models through Joint-Embedding Predictive Architectures DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 25

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no resolver link, observed 2026-08-02T22:21:33.203727Z

Source-reported events for the cited work

Unavailable: canonical work link unavailable.

source=pdf_text observed=2026-08-02T22:21:33.203727Z digest=sha256:3a3adbeb267e5af2704d602ccd0fae828b628453ff3f8eb345dad2e55987924d

Observation 8641dfd4-aebe-4672-9c04-a3bbe9a42a5d · inbound

In Search of Lost DNA Sequence Pretraining cites this paper.

In Search of Lost DNA Sequence Pretraining DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 40

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arxiv_id, observed 2026-05-10T08:53:03.604418Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

source=pdf_text observed=2026-05-10T08:52:47.515343Z digest=sha256:b09cf92f8e221742c2e26289978cfd21bc747b0267205d72b256925656b96952

Observation 781154fb-33c9-4fd7-900a-a6948c520d5a · inbound

Evaluating Post-hoc Explanations of the Transformer-based Genome Language Model DNABERT-2 cites this paper.

Evaluating Post-hoc Explanations of the Transformer-based Genome Language Model DNABERT-2 DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 31

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arxiv_id, observed 2026-05-09T22:34:07.319885Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

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Observation 0f381a55-dcf4-4edd-ac8f-038565affa18 · inbound

Set-Aggregated Genome Embeddings for Microbiome Abundance Prediction cites this paper.

Set-Aggregated Genome Embeddings for Microbiome Abundance Prediction DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 16

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arxiv_id, observed 2026-05-13T03:07:08.655044Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

source=arxiv_source observed=2026-05-13T03:06:39.232355Z digest=sha256:085cefa7b39bdffb230296cad96153c2dc346cbd521e0d8f89a903a245c51091

Observation 386ea257-d437-4f4b-8fba-85e2d3dbe1a1 · inbound

GGBound: A Genome-Grounded Agent for Microbial Life-Boundary Prediction cites this paper.

GGBound: A Genome-Grounded Agent for Microbial Life-Boundary Prediction DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 15

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verified exact
arxiv_id, observed 2026-05-15T02:13:30.434972Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

source=pdf_text observed=2026-05-15T02:12:05.190090Z digest=sha256:e3cba7af777a30dc85c83d99f5b522326e9e483b8a82d097d53a1fbbd7c90cda

Observation b5085feb-692b-4a27-802c-75ff4919adbd · inbound

TadA-Bench: A Million-Variant Benchmark for Future-Round Discovery Toward Agentic Protein Engineering cites this paper.

TadA-Bench: A Million-Variant Benchmark for Future-Round Discovery Toward Agentic Protein Engineering DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 99

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arxiv_id, observed 2026-06-28T20:12:37.859641Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

source=arxiv_source observed=2026-06-28T20:01:10.638647Z digest=sha256:6586fc1e889045fdf572ec90c32c8eea60bee13181319bfda4b0f0ae6545ab6f

Observation a931e25b-137b-4173-a238-d0e42cceefdd · inbound

Biological Reasoning-Informed Regression for Interpretable Regulatory DNA Activity Prediction cites this paper.

Biological Reasoning-Informed Regression for Interpretable Regulatory DNA Activity Prediction DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 44

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arxiv_id, observed 2026-07-02T22:17:26.272078Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

source=pdf_text observed=2026-06-27T19:01:43.892354Z digest=sha256:b89901683a4d21103c4bb163e8172c07b2267494a04f6bbf4072095e7104f5e4

Observation f9619132-339f-4b0b-bdb6-0131eb2f648e · inbound

Flexible Flows for Biological Sequence Design cites this paper.

Flexible Flows for Biological Sequence Design DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 9

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arxiv_id, observed 2026-07-03T03:57:38.577658Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

source=arxiv_source observed=2026-06-27T14:21:51.854183Z digest=sha256:003fcc2c99f4202a000efbfac732a7501ef09fec8058cab2c271bb2ade415645

Observation 2eeb9d5a-7574-4dad-b1c5-9b1cef876972 · inbound

How Post-Training Shapes Biological Reasoning Models cites this paper.

How Post-Training Shapes Biological Reasoning Models DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 36

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arxiv_id, observed 2026-07-01T07:55:31.056974Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

source=pdf_text observed=2026-07-01T07:48:31.110861Z digest=sha256:0f8eddedf75b1914d99a6a4d56d01da1f4d0edd924357b7c5b9973ba76b6bdb3

Observation 5fbac5d3-2744-45d5-b53c-3d21a2c7644f · inbound

DNA Language Models: An Assessment of Pre-Training for Fine-Tuning Tasks cites this paper.

DNA Language Models: An Assessment of Pre-Training for Fine-Tuning Tasks DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 13

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arxiv_id, observed 2026-06-30T03:34:13.167209Z

Source-reported events for the cited work

No event found in the named queried sources as of 2026-08-09T06:31:02.800959+00:00.

source=pdf_text observed=2026-06-30T03:32:20.018780Z digest=sha256:2fc49dec12f78a56882a046466315a4799b3f898533b0c3d0045d67100180625

Observation 5f819af3-be52-41d5-af98-ccd4d3ec50e6 · inbound

Deep and Probabilistic Models for Gene Regulatory Network Inference cites this paper.

Deep and Probabilistic Models for Gene Regulatory Network Inference DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 160

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no resolver link, observed 2026-08-01T21:34:24.389119Z

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Unavailable: canonical work link unavailable.

source=arxiv_source observed=2026-08-01T21:34:24.389119Z digest=sha256:de5f8dddd907970e37c5269a8057e1f9d2dc58502b38219e2dbd30beeac81ef2

Observation 7c860de4-861a-441c-a1d0-47819840e889 · inbound

Frozen but Not Always Accessible: A Representation Analysis of Genomic Language Models cites this paper.

Frozen but Not Always Accessible: A Representation Analysis of Genomic Language Models DNABERT-2: Efficient Foundation Model and Benchmark For Multi-Species Genome

Reference 55

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no resolver link, observed 2026-08-08T15:15:00.033761Z

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source=arxiv_source observed=2026-08-08T15:15:00.033761Z digest=sha256:4bfb0cd22e0f109468573921f7f73dc3c080b1dacb7c5639131ca2d524b826ef